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Family Cichlidae (Cichlids) - Sven Kullander

Family Cichlidae (Cichlids) - Sven Kullander

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Check List of the Freshwater Fishes of South and Central America<br />

Heros severus Heckel, 1840<br />

Heros severus Heckel, 1840: 362. Type locality: bei Marabitanas<br />

im Rio-negro. Syntypes: ?NMW 17354 (1), ?17638 (1), ?17656<br />

(1).<br />

Maximum length: 20 cm SL<br />

Distribution: South America: Orinoco River basin, in the upper<br />

Orinoco River drainage in Colombia and Venezuela; Amazon<br />

River basin, in the upper Negro River basin.<br />

Countries: Brazil, Colombia, Venezuela<br />

Heros spurius Heckel, 1840<br />

Heros spurius Heckel, 1840: 368. Type locality: Rio Guaporè ...<br />

Sümpfen. Syntypes: ?NMW 17656 (1), 17939 (1).<br />

Heros coryphaeus Heckel, 1840: 364. Type locality: Rio-Guaporé<br />

... Morästen um Matogrosso. Holotype: NMW 17358.<br />

Heros modestus Heckel, 1840: 366. Type locality: Rio Guaporé.<br />

Syntypes: NMW 17354 (1), ?92454 (1).<br />

Maximum length: 12 cm SL<br />

Distribution: South America: Amazon River basin, in the Guaporé<br />

River drainage, Brazil and Bolivia.<br />

Countries: Bolivia, Brazil<br />

HEROTILAPIA<br />

Herotilapia Pellegrin, 1904: 247. Type species: Heros multispinosus<br />

Günther, 1869. Type by monotypy. Gender: feminine.<br />

Herotilapia multispinosa (Günther, 1867)<br />

Heros multispinosus Günther, 1867: 601. Type locality: Lake of<br />

Managua. Holotype: BMNH 1865.7.20.34.<br />

Maximum length: 9 cm SL<br />

Distribution: Central America: Atlantic slope, from the Patuca<br />

River (Honduras) to the Matina River (Costa Rica); Pacific slope,<br />

from the Guasaule River (Nicaragua) to the Tempisque and Bebedero<br />

Rivers (Costa Rica).<br />

Countries: Costa Rica, Honduras, Nicaragua<br />

Remarks and references: Described in more detail in Günther<br />

(1868: 453, pl. 74, fig. 2). Reviewed by Bussing (1998: 351),<br />

with key, diagnosis, geographical distribution, and figures.<br />

Common names: Cholesca (Costa Rica), Mojarrita (Costa Rica),<br />

Zarzapala (Costa Rica)<br />

HOPLARCHUS<br />

Hoplarchus Kaup, 1860: 128. Type species: Hoplarchus pentacanthus<br />

Kaup, 1860. Type by subsequent designation. Gender: masculine.<br />

Hoplarchus psittacus (Heckel, 1840)<br />

Heros psittacus Heckel, 1840: 369. Type locality: Rio-negro,<br />

nördlich von Marabitanas am Fusse des Berges Cocui. Syntypes:<br />

?NMW 17641-42 (1, 1).<br />

Centrarchus cychla Jardine in Schomburgk, 1843: 157, pl. 11.<br />

Type locality: Rio Negro. No types known.<br />

Hoplarchus pentacanthus Kaup, 1860: 129, pl. 6 (fig. 1). Type<br />

locality: ? Südamerika. Holotype in ZSM, lost.<br />

Maximum length: 23.5 cm SL<br />

Distribution: South America: Amazon River basin, in the Negro,<br />

Preto da Eva, Urubu, and Jamari Rivers in Brazil; Orinoco River<br />

basin, in tributaries of the upper Orinoco River basin.<br />

Countries: Brazil, Venezuela<br />

HYPSELECARA<br />

Hypselecara <strong>Kullander</strong>, 1986: 232. Type species: Heros temporalis<br />

Günther, 1862. Type by original designation. Gender: feminine.<br />

Hypselecara coryphaenoides (Heckel, 1840)<br />

Heros coryphaenoides Heckel, 1840: 373, pl. 29 (fig. 9-12). Type<br />

locality: Rio-negro ... bei Maribitana. Holotype: NMW 17376.<br />

Heros niger Heckel, 1840: 375. Type locality: Rio-negro. Holotype:<br />

NMW 17358.<br />

Centrarchus niger Jardine in Schomburgk, 1843: 159, pl. 12. Type<br />

locality: Rio Negro. No types known.<br />

Cichlasoma arnoldi Ahl, 1936a: 138. Type locality: Amazonas.<br />

Holotype: ZMB 32382.<br />

Chuco axelrodi Fernández-Yépez, 1972: 15, fig. 16. Type locality:<br />

Aguaro River, Venezuela. Holotype: USNM 206097.<br />

Maximum length: 16 cm SL<br />

Distribution: South America: Amazon River basin, in the Negro,<br />

Trombetas, Tapajós, Maués, and Uatumã Rivers; Orinoco River<br />

basin in tributaries of the upper Orinoco River, and in the Aguaro<br />

River basin.<br />

Countries: Brazil, Venezuela<br />

Hypselecara temporalis (Günther, 1862)<br />

Heros temporalis Günther, 1862: 286. Type locality: No locality.<br />

Holotype: BMNH 1855.12.26.638 [ex Zool. Soc.].<br />

Acara (Heros) crassa Steindachner, 1875: 88, pl. 5. Type locality:<br />

Amazonenstrom bei Teffé, Tonantins, Cudajas, Coary, Villa<br />

bella und Obidos, im See Hyanuary und Saraca, so wie im Rio<br />

Hyutay. Syntypes: (several) NMW 17373-17374 (2) Codajás,<br />

33617 (1) Silves, 33618 (1) Villa Bella.<br />

Heros goeldii Boulenger, 1897: 298. Type locality: Upper Cunani<br />

River, French Guiana, south of the Oyapok River. Syntypes:<br />

BMNH 1897.7.17.9 (1), NMB 2488 (1).<br />

Cichlasoma Hellabrunni Ladiges, 1942: 199, fig. 1. Type locality:<br />

Gefangenschaftsnachzucht Hellabrunn (originally: entweder die<br />

Umgebung von Rockstone am Essequibo in Britisch-Guiana oder<br />

aber die Umgebung von Leticia. Lectotype: ZMH H70, designated<br />

by Ladiges et al. (1958: 164).<br />

Maximum length: 15 cm SL<br />

Distribution: South America: Amazon River basin, in the Ucayali<br />

and Amazon River drainages in Peru, the Amazon River drainage<br />

in Colombia, the Solimões-Amazon River in Brazil, east to Cametá,<br />

also rivers of Amapá State and the Oyapock River basin in<br />

Brazil.<br />

Countries: Brazil, Colombia, Peru<br />

Common names: Acará açai (Brazil), Bufurque (Peru), Bujurqui<br />

(Peru)<br />

HYPSOPHRYS<br />

Hypsophrys Agassiz, 1859: 408. Type species: Hypsophrys unimaculatus<br />

Agassiz, 1859. Type by monotypy. Gender: feminine.<br />

Copora Fernández-Yépez, 1969: [4]. Type species: Heros nicaraguensis<br />

Günther, 1864. Type species by original designation.<br />

Gender: feminine.<br />

Hypsophrys nicaraguensis (Günther, 1864)<br />

Hypsophrys unimaculatus Agassiz, 1859: 408. Type locality: Lake<br />

Nicaragua. Types in MCZ, not found.<br />

Heros nicaraguensis Günther, 1864c: 153. Type locality: Lake of<br />

Nicaragua. Holotype: BMNH 1867.9.23.37.<br />

Heros balteatus Gill in Gill & Bransford, 1877: 184. Type locality:<br />

Lake Nicaragua. Syntypes: BMNH 1905.3.27.3 [ex USNM] (1).<br />

Cichlasoma spilotum Meek, 1912: 73. Type locality: Victoria,<br />

Costa Rica. Holotype: FMNH 7686.<br />

Maximum length: 16.5 cm SL<br />

Distribution: Central America: Atlantic slope, from the San Juan<br />

drainage, including Lake Nicaragua, in Costa Rica and Nicaragua,<br />

to the Matina River drainage in Costa Rica.<br />

Countries: Costa Rica, Nicaragua<br />

Remarks and references: Species monograph by López (1974),<br />

well illustrated. <strong>Kullander</strong> & Hartel (1997) discussed the nomenclature<br />

of Hypsophrys unimaculatus. Reviewed by Bussing<br />

(1998: 354), with key, diagnosis, geographical distribution, and<br />

figures.<br />

637

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