An Updated Classification of the Recent Crustacea

otoidea on the recommendation of L. Holthuis
(1993a:87, and pers. comm.).
Superfamily Palaemonoidea
The family Euryrhynchidae Holthuis, 1950, was
added on the recommendation of Holthuis (pers.
comm.). The family Kakaducarididae was described
by A. J. Bruce (1993) as a subfamily of the
Palaeomonidae and is here treated as a family on
the recommendation of L. Holthuis (pers. comm.).
Superfamily Alpheoidea
Authorship of the family Ogyrididae remains credited
to Holthuis (1955). Although Hay and Shore
(1918) established the family Ogyridae, as noted by
M. Tavares (pers. comm.), L. Holthuis (pers.
comm.) points out that they based it on the type
genus Ogyris Stimpson, 1860, which is a junior
homonym of Ogyris Westwood and is thus invalid.
Stebbing (1914) proposed the replacement genus
Ogyrides, and thus the family name is Ogyrididae,
first used as such by Holthuis (1955). We have not
followed Christoffersen’s (1987) suggestion to
transfer the family Processidae to the Crangonoidea
or to combine the alpheoids and crangonoids and
pandaloids into one monophyletic taxon. Christoffersen
(1987) also proposed the new alpheoid families
Nauticarididae (to contain Nauticaris and Saron),
Alopidae (to contain Chorismus, Alope, and
Caridion), and Bythocarididae (to contain Bythocaris,
Cryptocheles, and Bathyhippolyte). We have
not followed these suggestions, nor have we recognized
the families Merhippolytidae and Thoridae
recognized by Christoffersen (e.g., Christoffersen
1998).
Christoffersen later (1987) also suggested the recognition
of the family Barbouridae (spelling corrected
to Barbouriidae by Christoffersen, 1990), to
include the genera Barbouria, Janicea, and Parhippolyte.
In his review of caridean shrimps of the Albatross
Philippine Expedition, Chace (1997), although
finding ‘‘no clear evidence to support the
superfamilial categories suggested by Christoffersen
(1987),’’ found ‘‘considerable reason to endorse his
[Christoffersen’s] establishment of the Barbouriidae.’’
Chace refrained from treating these genera
as Barbouriidae in that paper, but we have taken
that step here and recognize the Barbouriidae. Inclusion
of the family in the superfamily Alpheoida
is because of the similarities to hippolytids (all three
genera were formerly treated as members of the
Hippolytidae).
Superfamily Crangonoidea
As noted above, Christoffersen (1987) proposed the
family Barbouriidae for the genera Barbouria, Janicea,
and Parhippolyte and originally placed the
family in the superfamily Crangonoidea. We treat
it here as a member of the Alpheoidea because of
the similarities to the alpheoid family Hippolytidae
(see Chace, 1997:40).
Superfamily Pandaloidea
Christoffersen (1989) suggested a new classification
of this superfamily, wherein he proposed many significant
changes. Three new families were proposed
(Plesionikidae for the genus Plesionika, Heterocarpoididae
for the genus Heterocarpoides, and Dorodoteidae
for the genus Dorodotes). In addition,
the family Physetocarididae was removed from its
own superfamily and placed in the Pandaloidea,
and the family Heterocarpidae was recognized. No
diagnoses of the new taxa were provided (although
character states were given), and we have opted to
not recognize these changes for now.
INFRAORDER ASTACIDEA
Although we are not recognizing the ‘‘Macrura
Reptantia’’ as a suborder (see above), for the most
part, we have followed the admittedly conservative
classification of Holthuis (1991) for the superfamilies
and families of the Astacidea (see also Williams,
1988, for classification of commercially important
lobster families). Holthuis, who was at the
time dealing only with the marine lobsters and so
did not include the parastacoids and astacoids,
treated marine astacideans as belonging to a single
superfamily Nephropoidea containing two families,
Thaumastochelidae and Nephropidae. Our classification
differs only in the inclusion of the Enoplometopoidea
(see below) and Gylpheoidea, the latter
placed by Holthuis among the infraorder Palinura
(his Palinuridea). Scholtz (1999) recently reviewed
the freshwater crayfishes (Astacoidea and Parastacoidea)
and argued that they are members of a distinct
clade, Astacida, that is not closely related to
clawed lobsters. However, strong molecular evidence
suggests that clawed lobsters are indeed the
sister group to the astacids (Crandall et al., 2000).
Superfamily Glypheoidea
The primitive family Glypheidae (the only extant
family in the Glypheoidea) has been transferred to
the Astacidea as per the recommendations of Forest
and Saint Laurent (1989). The taxon name, credited
to Zittel in Bowman and Abele (1982), has
now been credited to the earlier usage by Winckler
(M. Hendrickx, pers. comm.), following the usage
in Glaessner (1969).
Superfamily Enoplometopoidea
The genus Enoplometopus was assigned its own superfamily
and family (Enoplometopoidea, Enoplometopidae)
by Saint Laurent (1988).
Superfamily Nephropoidea
Tshudy and Babcock (1997) examined fossil and
extant clawed lobsters and indicated that the family
Thaumastochelidae, at least as used previously,
may be paraphyletic. We have not taken the extra
step of deleting this family (which would result in
46 Contributions in Science, Number 39 Rationale