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An Updated Classification of the Recent Crustacea

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PREFACE<br />

For anyone with interests in a group <strong>of</strong> organisms<br />

as large and diverse as <strong>the</strong> <strong>Crustacea</strong>, it is difficult<br />

to grasp <strong>the</strong> enormity <strong>of</strong> <strong>the</strong> entire taxon at one<br />

time. Those who work on crustaceans usually specialize<br />

in only one small corner <strong>of</strong> <strong>the</strong> field. Even<br />

though I am sometimes considered a specialist on<br />

crabs, <strong>the</strong> truth is I can pr<strong>of</strong>ess some special knowledge<br />

about only a relatively few species in one or<br />

two families, with forays into o<strong>the</strong>r groups <strong>of</strong> crabs<br />

and o<strong>the</strong>r crustaceans. Crabs are but a small picture<br />

<strong>of</strong> <strong>the</strong> overall diversity <strong>of</strong> <strong>the</strong> <strong>Crustacea</strong>. They represent<br />

only one infraorder [Brachyura] within one<br />

order [Decapoda] within one superorder [Eucarida]<br />

within one subclass [Eumalacostraca] within one<br />

class [Malacostraca] <strong>of</strong> <strong>the</strong> six currently recognized<br />

classes <strong>of</strong> <strong>the</strong> <strong>Crustacea</strong> (as depicted herein). I am<br />

certain that this situation is similar for all o<strong>the</strong>r<br />

crustacean systematists, with <strong>the</strong> result that <strong>the</strong>re<br />

are no living specialists who can truly claim to have<br />

an in-depth understanding <strong>of</strong> <strong>the</strong> <strong>Crustacea</strong> as a<br />

whole.<br />

This volume is an attempt to provide <strong>the</strong> reader,<br />

whe<strong>the</strong>r a seasoned systematist or a beginning student,<br />

with a glimpse into <strong>the</strong> enormous variety <strong>of</strong><br />

extant crustaceans. The sheer number <strong>of</strong> categories<br />

that humans have constructed to contain and order<br />

this group is some indication <strong>of</strong> <strong>the</strong> incredible<br />

amount <strong>of</strong> morphological diversity <strong>the</strong>y exhibit.<br />

But this is only a small part <strong>of</strong> <strong>the</strong> overall picture.<br />

Even if one were to grasp <strong>the</strong> full range <strong>of</strong> taxonomic<br />

diversity as presented in this classification,<br />

such knowledge would shed no light on <strong>the</strong> actual<br />

biology <strong>of</strong> <strong>the</strong>se fascinating animals: <strong>the</strong>ir behavior,<br />

feeding, locomotion, reproduction; <strong>the</strong>ir relationships<br />

to o<strong>the</strong>r organisms; <strong>the</strong>ir adaptations to <strong>the</strong><br />

environment; and o<strong>the</strong>r facets <strong>of</strong> <strong>the</strong>ir existence<br />

that fall under <strong>the</strong> heading <strong>of</strong> biodiversity.<br />

By producing this volume we are attempting to<br />

update an existing classification, produced by Tom<br />

Bowman and Larry Abele (1982), in order to arrange<br />

and update <strong>the</strong> <strong>Crustacea</strong> collection <strong>of</strong> <strong>the</strong><br />

Natural History Museum <strong>of</strong> Los <strong>An</strong>geles County.<br />

This enormous and diverse collection contains an<br />

estimated four to five million specimens, making it<br />

<strong>the</strong> second largest collection <strong>of</strong> <strong>Crustacea</strong> in <strong>the</strong><br />

Americas. While undertaking this task, it occurred<br />

to us that o<strong>the</strong>rs might benefit from our efforts, and<br />

that perhaps a general update on <strong>the</strong> number and<br />

arrangement <strong>of</strong> <strong>the</strong> living crustacean families, along<br />

with an explanation <strong>of</strong> <strong>the</strong> systematic and classificatory<br />

changes suggested during <strong>the</strong> last two decades,<br />

might be a welcome addition to <strong>the</strong> literature.<br />

I hope this volume is seen as nothing more<br />

than <strong>the</strong> briefest <strong>of</strong> introductions into an understanding<br />

<strong>of</strong> crustaceans and that it might lead to<br />

fur<strong>the</strong>r work not only on <strong>the</strong> relationships among<br />

crustaceans but also toward understanding <strong>the</strong><br />

overall picture <strong>of</strong> crustacean biodiversity and natural<br />

history.<br />

Joel W. Martin<br />

June 2001<br />

Los <strong>An</strong>geles, California

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