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ICARDA annual report 2004

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accessions (Fig. 20) were sent to<br />

ICRISAT for DNA extraction and<br />

molecular analysis as per agreement<br />

in the Challenge Program.<br />

Fig. 20. Types of accessions from <strong>ICARDA</strong>’s chickpea collection<br />

chosen for inclusion in a global composite<br />

germplasm set that will be used to find valuable genes<br />

for crop breeding.<br />

<strong>ICARDA</strong>’s collection of nearly<br />

26,000 barley accessions is the second<br />

largest in the world. It includes<br />

15,500 accessions of locally-adapted<br />

landraces, a quarter of the global<br />

total, and 8700 accessions from<br />

CWANA, the largest collection of<br />

landraces from that region. The<br />

Center’s 1800 accessions of H. vulgare<br />

ssp. spontaneum represent its<br />

full geographical range.<br />

Using GIS, researchers generated<br />

detailed agroclimatological<br />

information, based on 67 variables,<br />

for accessions with collection-site<br />

Fig. 21. Types of accessions chosen by <strong>ICARDA</strong> from its<br />

barley collection for inclusion in a global composite<br />

germplasm set that will be used to find valuable genes<br />

for crop breeding.<br />

coordinates: 72% of the landrace<br />

accessions and 52% of the wild progenitor<br />

accessions. Landrace and<br />

wild barley accessions were selected<br />

for the composite<br />

set by analyzing this<br />

information using a<br />

two-step cluster<br />

analysis. This produced<br />

260 clusters,<br />

from which<br />

researchers selected<br />

accessions from different<br />

geographical<br />

areas. Improved<br />

germplasm was<br />

selected for the barley<br />

composite set using<br />

passport information,<br />

including pedigrees,<br />

to ensure that the<br />

most common parental varieties<br />

and lines were represented.<br />

<strong>ICARDA</strong> barley breeders also chose<br />

improved drought-tolerant<br />

germplasm for inclusion.<br />

The final barley composite set<br />

(Fig. 21) consisted of 445 accessions<br />

of H. vulgare ssp. spontaneum (15%<br />

of the total), 1935 landrace accessions<br />

(65%), and improved<br />

germplasm (20%). The improved<br />

germplasm category includes cultivars,<br />

unfinished breeders’ materials,<br />

and genetic stocks, representing<br />

13%, 6%, and 1% of the composite<br />

barley set, respectively.<br />

Of the barley wild<br />

progenitor accessions,<br />

65% are original<br />

accessions collected<br />

during GRU missions.<br />

Hyper-arid, arid and<br />

semi-arid collection<br />

sites represent 1%,<br />

20%, and 63% of the<br />

total, respectively.<br />

These accessions originate<br />

from 20 coun-<br />

tries and their collection<br />

sites belong to 58<br />

ecological clusters.<br />

Of the landraces,<br />

Theme 3<br />

20% are original materials collected<br />

by <strong>ICARDA</strong>. Hyper-arid, arid and<br />

semi-arid collection sites represent<br />

3%, 33%, and 43% of the total,<br />

respectively. The landrace set originates<br />

from 85 countries, and 78%<br />

are from CWANA. Collection sites<br />

belong to 255 ecological clusters.<br />

Exploring variation in<br />

barley’s photothermal<br />

response to meet the<br />

challenges of climate<br />

change<br />

Global warming is expected to<br />

reduce grain yields in rainfed farming<br />

systems in CWANA’s arid and<br />

semi-arid regions. To develop crop<br />

varieties able to cope with climate<br />

change, breeders are working to<br />

optimize the timing of the crop’s<br />

growing season in relation to temperatures<br />

and the amount of rainfall<br />

expected. This requires the use<br />

of traits associated with a crop’s<br />

photothermal response. To identify<br />

these traits, researchers from<br />

<strong>ICARDA</strong> and Germany’s Institute<br />

of Genetics and Plant Breeding are<br />

evaluating <strong>ICARDA</strong>’s germplasm<br />

collection as part of a project funded<br />

by the German Federal Ministry<br />

for Economic Cooperation and<br />

Development (BMZ) and GTZ.<br />

The speed with which cereals<br />

reach the heading stage is controlled<br />

by three separate genetic<br />

mechanisms which determine (i)<br />

response to temperature over a<br />

period of time (earliness per se); (ii)<br />

response to day length (photoperiod<br />

sensitivity); and (iii) response to<br />

low temperatures at the initial<br />

stages of plant development (vernalization<br />

sensitivity). Researchers<br />

studied these responses in 77<br />

improved barley lines and cultivars<br />

from <strong>ICARDA</strong> breeders’ collections,<br />

and in 115 landraces and 35<br />

wild barley (Hordeum vulgare ssp.<br />

<strong>ICARDA</strong> Annual Report <strong>2004</strong><br />

53

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