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Chapter 6 Biotic Resources • 99Figure 6.1 • The growth, or sustainable yield, curve.been driven extinct (the origin in Figure 6.1). In between these twopoints, things get interesting.The growth, or sustainable yield, curve for a renewable natural resourceindicates the increase in stock over one time period for any given stock.The y-axis can measure growth or harvest, that is, flows from or to the existingstock depicted on the x-axis. Any harvest up to the total stock is theoreticallypossible. A harvest at any point on the sustainable yield curve,such as S, is just equal to the growth in the stock and hence has no net impacton stocks. Harvests above the sustainable yield curve deplete the resource,and harvests below the curve lead to an increase in the stock, asindicated by the arrows. For example, a harvest at R will reduce the stockto S′′, and a harvest at S′ will allow the stock to increase to R′′. MVP representsthe minimum viable population, which is the level below which aspecies or stock cannot sustain itself even in the absence of harvest. Thephrase critical depensation refers to the spontaneous decline of a populationor ecosystem that has fallen below the minimum viable population orsize. 5 K represents carrying capacity. The graph suggests extremely rapidgrowth rates—about 30% per time period at maximum sustainable yield(MSY), which is the average maximum catch that can be removed underexisting environmental conditions over an indefinite period without causingthe stock to be depleted, assuming that removals and natural mortalityare balanced by stable recruitment and growth. While this may be appro-5 The first edition of this textbook used the term critical depensation level instead of minimumviable population. Critical depensation is useful when referring to ecosystems that cannot sustainthemselves when stocks fall below a certain level, since it makes little sense to talk about size ofan ecosystem as a population. However, minimum viable population, in general, is a more intuitivelyobvious phrase.

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