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modern variation and evolutionary change in the hominin eye orbit

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differences <strong>in</strong> measurement technique will not confound <strong>the</strong> <strong>in</strong>vestigation of populationaff<strong>in</strong>ities. Though <strong>the</strong>se two techniques yield consistent results, because a differentresearcher collected <strong>the</strong>se <strong>orbit</strong>al volume data as part of a separate study, <strong>the</strong> degree of<strong>in</strong>terobserver error cannot be known.In addition to this <strong>orbit</strong>al volume measurement recorded <strong>in</strong> milliliters, a separate<strong>in</strong>dicator of <strong>orbit</strong>al size is used <strong>in</strong> <strong>the</strong> analysis of <strong>evolutionary</strong> <strong>change</strong> <strong>in</strong> <strong>orbit</strong>almorphology s<strong>in</strong>ce <strong>the</strong> Upper Paleolithic <strong>in</strong>vestigated <strong>in</strong> chapter 4. This provides anadditional estimate that enables <strong>the</strong> <strong>in</strong>clusion of more <strong>in</strong>dividuals that would o<strong>the</strong>rwise beexcluded due to poor preservation of <strong>the</strong> th<strong>in</strong> bones that make up <strong>the</strong> <strong>in</strong>terior <strong>orbit</strong>.This <strong>orbit</strong>al size variable is derived from <strong>the</strong> geometric mean of <strong>orbit</strong>al height,<strong>orbit</strong>al breadth, <strong>and</strong> <strong>orbit</strong>al depth (Height * Width * Depth)·³³³³³ follow<strong>in</strong>g <strong>the</strong> methods ofGonzález-José et al. (2005). The geometric mean of <strong>the</strong>se three variables provides anestimate of volume, which is <strong>the</strong> same as that of a rectangular prism with side lengthsequal to each measure of <strong>orbit</strong>al depth, width, <strong>and</strong> height. Though <strong>the</strong> <strong>orbit</strong> is more pearshapedthan cuboid, this provides an estimate of its volume when damage to <strong>the</strong> fragilebone of its <strong>in</strong>ternal structure prohibits accurate measure <strong>in</strong> milliliters.In addition to <strong>the</strong>se cont<strong>in</strong>uous traits of <strong>the</strong> face <strong>and</strong> cranium, o<strong>the</strong>r quantitative<strong>and</strong> categorical variables such as age, sex, ancestry, species, <strong>and</strong> time are used foraddress<strong>in</strong>g questions <strong>and</strong> test<strong>in</strong>g hypo<strong>the</strong>ses relat<strong>in</strong>g to <strong>orbit</strong>al development, <strong>in</strong>terspecific<strong>variation</strong>, sex differences, <strong>and</strong> how much <strong>change</strong> is characteristic of this feature throughtime. In most cases sex was listed <strong>in</strong> <strong>the</strong> museum database or written on <strong>in</strong>dividualskulls, though when not available it was estimated us<strong>in</strong>g different skeletal <strong>in</strong>dicators afterBurns (1999), <strong>and</strong> was marked <strong>in</strong>determ<strong>in</strong>ate if a clear dist<strong>in</strong>ction could not be made. In33

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