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modern variation and evolutionary change in the hominin eye orbit

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upward of 5 times for 23 different skulls at different museums over <strong>the</strong> course of 4 yearsof data collection. In each case dimensions were recorded from a r<strong>and</strong>omly selectedskull, which was <strong>the</strong>n set aside until later <strong>in</strong> <strong>the</strong> day or for <strong>the</strong> follow<strong>in</strong>g day to ensurethat repeated measures wouldn’t be biased by consecutive quantification. Measurementerror was assessed follow<strong>in</strong>g <strong>the</strong> method described <strong>in</strong> White (2000), page 307.Variables of <strong>the</strong> cranium <strong>and</strong> face were found to be slightly more consistent,deviat<strong>in</strong>g only 0.56% from <strong>the</strong> average of each measurement. By comparison, variablesof <strong>the</strong> <strong>orbit</strong> were found to diverge by 1.07% on average. Though most <strong>orbit</strong>al variablesare as accurate or <strong>in</strong> some cases more accurate than those taken from o<strong>the</strong>r features of <strong>the</strong>cranium <strong>and</strong> face, <strong>in</strong>ter<strong>orbit</strong>al breadth, <strong>orbit</strong>al volume, <strong>and</strong> <strong>the</strong> distance from basion to<strong>orbit</strong>ale have measurement errors above 1%, which acts to <strong>in</strong>crease <strong>the</strong> average error for<strong>the</strong>se <strong>orbit</strong>al variables (Tables 2.4 & 2.5).Seeds used to estimate volume of <strong>the</strong> <strong>orbit</strong> were sifted to ensure that each wasroughly <strong>the</strong> same size, <strong>and</strong> <strong>the</strong> same seeds were used at each museum where data werecollected. Additional precautions were taken to ensure that <strong>the</strong> measurement techniquewas consistent throughout, as it was recognized prior to beg<strong>in</strong>n<strong>in</strong>g data collection thatsome error is <strong>in</strong>herent <strong>in</strong> volumetric estimations of <strong>the</strong> <strong>orbit</strong> us<strong>in</strong>g seed, partially due to<strong>the</strong> less well def<strong>in</strong>ed border <strong>in</strong> <strong>the</strong> upper corner of <strong>the</strong> medial <strong>orbit</strong>al marg<strong>in</strong> (Schultz,1940). Even with <strong>the</strong>se precautions <strong>and</strong> <strong>the</strong> meticulousness with which <strong>orbit</strong>al volumewas taken, measurement of this trait is found to be <strong>the</strong> least repeatable of all variables,with an average difference from <strong>the</strong> mean measurement of 2.97%.35

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