DEPARTMENT OF ETHOLOGY AND ECOLOGY Head of ... - DPZ

Farbprofil: Generisches Offset-Separations-Profil 

Schwarz 150 lpi bei 45 Grad 

Department of Ethology and Ecology 

DEPARTMENT OF ETHOLOGY AND ECOLOGY 

Head of department: PD Dr. Peter M. Kappeler 

General research objectives 

Within the wide fields of behaviour and ecology, our current research activities focus 

on two central topics: primate social systems and community ecology. We conduct 

comparative field studies on these topics with lemurs in Madagascar, tamarins and 

titi monkeys in Peru, and baboons in Eritrea. Our field projects are multidisciplinary 

and involve a number of internal and external collaborations to study behavioural, 

endocrinological and genetic questions. By addressing similar questions in these different 

projects within a common theoretical framework, we create a basis for identifying 

basic processes and mechanisms in the evolution of primate behavior and ecology. 

These and other results from completed studies of langurs in Nepal and titi monkeys 

in Brasil are being used to test hypotheses on the evolution of primate communities 

and develop recommendations for conservation projects. 

Structure of department 

By the end of 1998, the department consisted of the provisional head of department, 

one postdoctoral scientist, two PhD students financed by the DPZ, one PhD 

student financed by a grant from the DFG, and one technician. A postdoctoral scientist 

financed by a grant from the DFG (part time position), four other postdoctoral 

scientists, three PhD students and several diploma students also worked in the department. 

In Madagascar, two PhD students and three diploma students are associated 

with the department; the field station in Peru is headed by a Peruvian diploma 

student. 

During the past year, several members of our department graduated or received 

additional degrees. Julia Ostner, Bärbel Häuser and Léon Razafimanantsoa completed 

their diploma degrees. After completing her diploma thesis, Cornelia Kraus 

was admitted to the doctoral program at Bielefeld University. Oliver Schülke, 

Manfred Eberle and Julia Ostner began their graduate studies in our department, 

and Christoph Knogge received his PhD from Bielefeld University. Stefanie Heiduck 

received the Dissertation Award of the Primate Center and Eckhard W. Heymann 

and Peter M. Kappeler obtained postdoctoral degrees in primatology and zoology, respectively. 

The part time positions of Carola Borries and Andreas Koenig could not 

be extended. 

Scientists 

Dr. Carola Borries (-31.07.98) 

Dr. Stefanie Heiduck 

PD Dr. Eckhard W. Heymann 

Dr. Andreas Koenig (-31.07.98) 

Dr. Dietmar Zinner 

Technician 

Ulrike Walbaum 

PhD Students 

Manfred Eberle 

Joanna Fietz 

Primate Report 54, July 1999 51 

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Dienstag, 7. September 1999 07:30:02 

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PhD Students 

Christoph Knogge 

Julia Ostner 

Rodin Rasoloarison 

Lèonard Razafimanantsoa 

Oliver Schülke 

Dorothea Schwab 

Barbara Wimmer 

Undergraduate Students 

Alexandra Dill 

Nikolaus von Engelhardt 

Andreas Hapke 

Bärbel Häuser 


Sonja Köhler 

Aina Rasamoelina 

Inga Timmann 

Emerita Tirado 

Trainees 

Katrin Böger 

Kathrin Dausmann 

Frederike Fellenberg 

Lisa Hoch 

Julia Küstner 

Pia Weidlich 

Andrea Weidt 

Nutrition and Seasonal Reproduction in Hanuman Langurs: Seasonal variation of 

energy intake and energy expenditure leads to strong variation in the physical condition 

of females (–•–). An above-average physical condition is reached first in July and 

coincides with the period of conceptions (bar diagram; values indicate the number of 

conceptions observed). The underlying hormonal changes are currently being investigated 

by the working group “Comparative Endocrinology”. 

52 Primate Report 54, July 1999 




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Progress during period under report 

Results 

In the past year, we made important progress on a number of fronts. In Madagascar, 

Oliver Schülke succeeded in capturing and radiocollaring fork-marked lemurs, 

so that it is now possible to initiate a field study of this intriguing lemur. Julia 

Ostner developed the necessary methodology for her study of the socio-endocrinology 

of redfronted lemurs. Barbara Wimmer developed the genetic tools necessary for the 

analyses of kin relations in several lemur species. Preliminary results of Rodin 

Rasoloarison's thesis work indicate the existence of at least four species of mouse lemurs 

that are new to science. 

Scent marking events / hour 

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p < 0.01 


Male Female 

Different aspects of olfactory communication were addressed in coordinated comparative 

studies in tamarins and lemurs. Moustached tamarins (left graph) show a pronounced 

sexual dimorphism in favour of females with regard to scent marking (from: 

Heymann, E.W. (1998): Behav. Ecol. Sociobiol. 43:40). Latency of reaction of male 

and female recipients to three different types of scent marks in Lemur catta (right 

hand graph). Mean latencies (+SE) between deposition of female anogenital (Female 

AG), male anogenital (Male AG) and male antebrachial (Male AB) scent marks and 

first reaction with the same signal in male (black bars) and female (hatched bars) recipients 

are shown. The type of scent mark, but not sex of recipient, significantly influences 

mean latency. In a two-way ANOVA, the interaction (type x sex of recipient) was 

not significant (from: Kappeler, P.M. (1998): Behav. Ecol. Sociobiol. 42:411). 

Considerable progress was also made in the research carried out in Peru. Emerita 

Tirado succeeded in habituating red titi monkeys, Callicebus cupreus, so that a new 

comparative study of their behavioral ecology can now be initiated. The comparisons 

with the completed studies by Stefanie Heiduck on Brazilian masked titi monkeys, 

Callicebus personatus, and the ongoing studies on moustached and saddle-back 

tamarins (Saguinus mystax, Saguinus fuscicollis) provide an opportunity to elucidate 

the relative importance of ecological and phylogenetic factors in the evolution of 

specific socio-ecological adaptations. Christoph Knogge and Emerita Tirado finished 





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their theses. Their results detailed the role of tamarins in seed dispersal and, thus, 

forest regeneration, and provide a basis for further specific studies of this aspect of 

ecology. 

In Eritrea, Dietmar Zinner completed a nationwide survey of hamadryas baboons 

and grivet monkeys. This information will be useful for local conservation activities. 

In addition, the classification of eco-zones was also completed, so that their covariation 

with primate distribution and abundance can now be examined. During this 

phase of the study significant methodological progress has been made in terms of an 

integration of ecological, demographic and socio-economic information in a GIS. All 

this experience will be very useful in future conservation projects of the DPZ in 

other parts of the world. 

External collaborations 

Despite the ongoing uncertainty of planning for the department, new external 

cooperations were established and existing ones extended. 

• A collaboration contract with the Parc Botanique et Zoologique de Tsimbazaza of 

Madagascar provides a framework for future collaborations. The first project will 

focus on the anatomy of the visual system of lemurs. 

• A collaboration with the Laboratoire d'Ecologie de MNHN à Paris (Dr. M. Perret) 

will allow the exchange of doctoral students to integrate our studies of reproduction 

and physiology in captivity and the field. 

• In a joint project with the MPI for Brain Research (PD Dr. L. Peichl), the anatomy 

of the visual system of lemurs with different activity patterns is being studied. 

• In collaboration with Prof. Dr. H. Erkert (University Tübingen) we now use actometers 

to register the activity of cathemeral lemurs. 

• The existing cooperation with the Universidad Nacional Mayor de San Marcos in 

Lima (F. Encarnación and R. Aquino) on the Biogeography and Conservation of 

Amazonian primates was intensified. 

• The cooperation between the Dept. of Landuse and Conservation of the University 

of Applied Science of Eberswalde and the University Autónoma de Madrid contributed 

strongly to the success of the hamadryas baboon project in Eritrea. Also 

the collaboration with the University of Asmara made considerable progress. During 

the next field campaign Eritrean students will participate for the first time in 

the hamadryas baboon project. Additionally, members of European primate research 

groups are becoming interested in a collaboration, e.g. from the University 

of Antwerp, Belgium. 

Field stations 

Work in Madagascar continued to make progress. The research station was 

partly remodeled and supplied with a new solar panel. Another 20 ha grid system 

was also established, further improving our working conditions. This increased our 

study area to more than 120 ha with about 80 km of trails. The station continued to 

be used by numerous external users and DPZ staff. Two field courses of the Universities 

in Antananarivo and Würzburg took place at our station and two film teams 





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visited the station to film the lemurs and their researchers. At two sites, temporary 

field camps have been established in Eritrea, one in the central highlands and a second 

one in the coastal lowlands. Habituation of hamadryas baboons at the sites has 

just begun. The field station in Peru (Estación Biológica Quebrada Blanco) was used 

more intensively by external users than in the years before. 

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AoS 

home-ranges 

Fc Fs Wc Ws Bu Gr Bs Ar Op Ot Cl Sh 

Vegetation structure in hamadryas baboon (Papio h. hamadryas) home-ranges in 

eastern and central Eritrea (N=40; means ± SE) and for comparison, the vegetation 

structure of the entire area of survey (AoS, 28.000 km 2 , *p40% coverage); 

Fs = medium closed forest (>2 layers, 10–40% coverage); Wc = closed woodland 

(1-2 layers, >40% coverage); Ws = medium closed woodland (1-2 layers, 10-40% coverage); 

Bu = bush- and shrubland; Gr = grassland; Bs = barren soil; Ar = agricultural 

areas; Op = Opuntia ficus-indica; Ot = others (settlements, roads, open inland water); 

Cl = cloud cover (not classified); Sh = shades from steep mountains (not classified). 

Events 

Madagascar was the focus of three scientific highlights during last year. Our department 

was represented by 9 active participants and numerous presentations at 

the Congress of the International Primatological Society in Antananarivo. Together 





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with more than 20 colleagues from North and South America, England and Italy, 

several researchers from DPZ took the opportunity to visit our field station in 

Kirindy for the first time. A few weeks earlier, the third Kirindy Symposium took 

place at DPZ. This symposium had more than 50 participants from Germany and 

abroad who reported on their work carried out over the last two years in Kirindy. 

Finally, we were all very happy that Rodin Rasoloarison and Léon Razafimanantsoa 

had the opportunity to leave Madagascar for the first time and to carry out research 

visits to DPZ as well as in Paris, Berlin and Chicago and to participate at the Kirindy 

Symposium. 

Projects and collaborators 

(I: internal cooperation, E: external cooperation; A: project completed, L: current project) 

Projects and collaborators of the 


Conservation biology of Malagasy lemurs 

B. RAKOTOSAMIMANANA (Univ. Madagascar, Antananarivo), 

J.U. GANZHORN (Institute of Zoology, Univ. Hamburg), S. GOODMAN 

(FMNH, Chicago & WWF, Madagascar), K. HODGES (Dept. of Reproductive 

Biology, DPZ), H. ZISCHLER (Working Group Primate Genetics, DPZ), 

P.M. KAPPELER 

Biogeography and conservation status of Amazon primates 

R. AQUINO, F. ENCARNACIÓN (Univ. Nacional Mayos San Marcos, 

Lima), E.W. HEYMANN 

Population genetics of hamadryas baboons 

A. HAPKE, H. ZISCHLER (Working Group Primate Genetics, DPZ), 

D. ZINNER 

Distribution patterns of primates in Eritrea 

F. PELÁEZ (Univ. Autónoma de Madrid, E), F. TORKLER, (Univ. 

Eberswalde), MINSTRY OF AGRICULTURE (Asmara, Eritrea), 

D. ZINNER 

Human impact on Nepalese forests and Hanuman langurs 

A. KOENIG, C. BORRIES, K. WESCHE (Dept. of Geography, 

Univ. Marburg) 

Taxonomy and biogeography of mouse lemurs 

R. RASOLOARISON (Univ. Madagascar, Antananarivo and Dept. of Ethology 

and Ecology, DPZ), S. GOODMAN (FMNH, Chicago & WWF, Madagascar), 

A. YODER (Northwestern Univ., USA), J.U. GANZHORN (Institute 

of Zoology, Univ. Hamburg), B. RAKOTOSAMIMANANA (Univ. 

Madagascar, Antananarivo), P.M. KAPPELER 

I,E L 

E L 

I L 

E A 

E A 

E L 





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Lemurs and forest regeneration in Western Madagascar 

J.U. GANZHORN (Institute of Zoology, Univ. Hamburg), E. RAKOTOVAO 

(CFPF, Morondava, Madagascar), J. FIETZ, D. SCHWAB, D. ZINNER 

E A 

Seed dispersal by tamarins, secondary dispersion and seed 

predation in Parkia 

M. FELDMANN (Dept. of Biology, Univ. Göttingen), E.W. HEYMANN 

Adaptations of the visual system of diurnal and nocturnal lemurs 

E L 

L. PEICHL (MPI for Brain Research, Frankfurt), A. RANDRIANJAFY 

(Parc Botanique et Zoologique de Tsimbazaza, Madagascar), 

P.M. KAPPELER 

E L 

Color vision and foraging strategies of tamarins 

H.M. BUCHANAN-SMITH (Univ. Stirling, GB), N. MUNDY (Univ. 

Oxford, GB), E.W. HEYMANN 

E L 

Genetic analysis of mating systems of polygynous lemurs 

B. WIMMER, D. TAUTZ (Univ. München), P.M. KAPPELER 

Socio-endocrinology of group-living lemurs 

E L 

M. HEISTERMANN (Dept. of Reproductive Biology, DPZ), J. OSTNER, 

N. VON ENGELHARDT, C. KRAUS, P.M. KAPPELER 

I L 

Proximate regulation and genetic consequences of the mating 

system of polyandric tamarins 

E.W. HEYMANN, U.-R. BÖHLE, C. SCHWIEGH (Working Group Primate 

Genetics, DPZ), M. HEISTERMANN, K. HODGES (Dept. of Reproductive 

Biology, DPZ) 

I L 

Olfactory communication in callitrichids 

B. HÄUSER, R. FRANKE (Univ. Gießen), E.W. HEYMANN 

Population ecology of Cheirogaleus medius 

E A 

J. FIETZ , J.U. GANZHORN (Institute of Zoology, Univ. Hamburg), 

U.-R. BÖHLE, C. SCHWIEGH (Working Group Primate Genetics, DPZ) 

I,E L 

Mating and reproductive strategies of male Hanuman langurs 

K. LAUNHARDT (Institute of Zoology and Anthropology, Univ. Göttingen), 

C. BORRIES, J.T. EPPLEN, C. EPPLEN (Molecular Human Genetics, 

Ruhr-Univ. Bochum), P. WINKLER (Tropical Centre, Univ. Göttingen) 

Ecology of male-female relations in Hamadryas baboons 

D. ZINNER, F. PELÁEZ, (Univ. Autónoma de Madrid, Spain), I. LEIN 

FELDER (Univ. Antwerp, B), J. SHOSHANI (Dept. of Biology, Univ. 

Asmara, Eritrea) 

E A 

E L 





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Sexual swellings of female Hamadryas baboons 

D. ZINNER, T. DESCHNER (MPI Evolutionary Anthropology, Leipzig) 

Social grooming in Hamadryas baboons 

E A 

D. ZINNER, I. TIMMANN (Institute of Zoology, Univ. Hamburg) 

Species concepts 

E L 

S. KÖHLER, H. ZISCHLER (Working Group Primate Genetics, DPZ), 

D. ZINNER 

I L 

Helper system of Cebuella pygmaea 

P. SOINI (Instituto de Investigaciones de la Amazonía Peruana, Iquitos, 

Peru), E.W. HEYMANN 

E L 

Activity of cathemeral lemurs 

H. ERKERT (Univ. Tübingen), P.M. KAPPELER, J. OSTNER 

Physiology and behaviour of mouse lemurs 

E L 

M. PERRET (MNHN, Laboratoire d'Ecologie, Brunoy, Frankreich), 

M. EBERLE, P.M. KAPPELER 

E L 

Acoustic communication in lemurs 

C. FICHTEL, K. HAMMERSCHMIDT (Dept. of Neurobiology, DPZ), 

P.M. KAPPELER 

Locomotor behavior of Hanuman langurs 

J. NIKOLEI (Institute of Anthropology und Human Biology, Free Univ. 

Berlin), C. BORRIES, A. KOENIG 

Feeding ecology of sympatric tamarins 

A.C. SMITH, H.O. BOX (Univ. Reading, GB), C. KNOGGE, E.W. 

HEYMANN 

I L 

E A 

E L 

Behavioural ecology and evolution of interspecific associations 

in callitrichines 

H.M. BUCHANAN-SMITH (Univ. Stirling, GB), E.W. HEYMANN 

Interspecific associations between cheirogaleids and endemic 

rodents in Madagascar 

E L 

D. SCHWAB, J.U. GANZHORN (Institute of Zoology, Univ. Hamburg) 

Costs of infant carrying in cotton-top tamarins 

E L 

S. SANCHEZ, F. PELÁEZ (Univ. Autonóma de Madrid, E), W. KAUM- 

ANNS (Dept. of Veterinary Medicine and Primate Husbandry, DPZ), 

E.W. HEYMANN 

E, I A 





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Phylogeny of New World primates 

S. SINGER, H. ZISCHLER, J. SCHMITZ (Working Group Primate 

Genetics, DPZ), E.W. HEYMANN 

Body mass of Callimico 

F. ENCARNACIÓN (Univ. Nacional Mayos San Marcos, Lima), 

E.W. HEYMANN 

Feeding ecology of Hanuman langurs 

A. KOENIG, C. BORRIES, M.K. CHALISE (Natural History Society 

of Nepal, Kathmandu), K. WESCHE (Dept. of Geography, Univ. 

Marburg), J.U. GANZHORN (Institute of Zoology, Univ. Hamburg), 

J. NIKOLEI (Institute for Anthropology and Human Biology, 

Free Univ. Berlin) 

Scientific contributions 

Habilitation theses 

I L 

E A 

E A 

HEYMANN, E.W.: Aspekte der Verhaltensbiologie von Schnurrbarttamarinen, 

Saguinus mystax (Callitrichinae; Cebidae; Primates) in ökologischer, evolutionärer 

und phylogenetischer Perspektive. University Gießen. 

KAPPELER, P.M.: Konvergenz und die soziale Evolution madagassischer Lemuren. 

University Würzburg. 

Doctoral theses 

KNOGGE, C.: Tier-Pflanze-Interaktionen im Amazonas-Regenwald: Samenausbreitung 

durch die sympatrischen Tamarinarten Saguinus mystax und Saguinus 

fuscicollis (Callitrichinae, Primates). University Bielefeld (1998). 

Diploma theses 


FRANKE, R.: Vergleichende Untersuchungen des Markierverhaltens von fünf Krallenaffenarten 

(Familie Callitrichidae) in Gefangenschaft. University Gießen (1998). 

HÄUSER, B.: Die Struktur der olfaktorischen Kommunikation in Familiengruppen 

von Lisztaffen, Saguinus oedipus (Primates: Plathyrrhini: Callitrichidae), in Gefangenschaft. 

University Gießen (1998). 

OSTNER, J.: Aktivitätsrhythmus, Sozial- und Paarungssystem einer kathemeralen 

Lemurenart (Eulemur fulvus rufus). Institute of Zoology II, Georg-August-University 

Göttingen (1998). 





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RAZAFIMANANTSOA, L.: Contribution à l'étude de locomotion de deux lèmuriens 

sympatriques: Propithecus verreauxi et Eulemur fulvus. Université de Madagascar 

Antananarivo (1998). 

TIRADO HERRERA, E.R.: Germinación de semillas dispersadas por Saguinus 

mystax y Saguinus fuscicollis (Callitrichidae), en comparación con semillas no 

dispersadas en Quebrada Blanco, Loreto, Perú. Facultad de Ciencias Biológicas, 

Universidad Nacional de la Amazonía Peruana (1998). 

Congress contributions 


11th Annual Meeting of the German Society for Tropical Ecology, Bielefeld, 20.- 

22.02.98, 

KNOGGE, C., HEYMANN, E.W.: Effektive Samenausbreitung durch Tamarine: der 

Einfluß der Darmpassage auf das Keimverhalten ausgebreiteter Samen. 

HEIDUCK, S.: Nutzung sekundärer Vegetation durch schwarzköpfige Springaffen 

(Callicebus personatus melanochir). 

HEYMANN, E.W.: Verhaltensökologische Anpassungen an die Nutzung einer 

"schwerverdaulichen" Nahrung: tageszeitliche Muster des Exsudatkonsums bei 

Saguinus mystax. 

Workshop "Diet, Foraging Behavior and Time-Budgets in Nonhuman Primates", European 

federation of Primatology, Paimpont (Frankreich), 16.-18.04.98, HEYMANN, 

E.W.: When to eat exudates - feeding itineraries in wild Saguinus mystax. 

3rd. Kirindy Symposium, DPZ, Göttingen, 10.07.98, 

FIETZ, J.: Social system and population ecology of the fat-tailed dwarf lemur 

(Cheirogaleus medius). 

OSTNER, J.: Activity period and lemur social systems - special relationships in the 

cathemeral Eulemur fulvus rufus? 

SCHÜLKE, O.: What you always wanted to know about Phaner (Phaner furcifer), 

but never dared to ask. 

SCHWAB, D.: Interspecific competition of three sympatric lemurs (Microcebus 

myoxinus, Microcebus murinus and Cheirogaleus medius) in Kirindy forest (West 

Madagascar). 

7th International Behavioral Ecology Congress, Pacific Grove, CA, USA., 27.07.- 

01.08.98, 

BORRIES, C., LAUNHARDT, K.: Infanticide is in accordance with sexual selection 

theory - DNA evidence for Hanuman langurs. 

KOENIG, A., BORRIES, C.: Costs of sociality in Hanuman langurs. 

OSTNER, J., KAPPELER, P.M.: The impact of cathemerality on the social system of 

wild red-fronted lemurs. 

SCHÜLKE, O.: The monopolization potential of langur males - is energy gain decisive? 

XVIIth Congress of the International Primatological Society, Antananarivo, Madagascar, 

09.-14.08.98, 





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DONATI, G., LUNARDINI, A., KAPPELER, P.M.: Cathemeral activity of Eulemur 

fulvus rufus in the Kirindy forest. 

EBERLE, M., KAPPELER, P.M.: Reproductive synchrony in lesser mouse lemurs 

(Microcebus murinus). 

HEYMANN, E.W.: Mixed-species troops in Saguinus and Cercopithecus - a comparison. 

FIETZ, J.: Monogamy in a nocturnal lemur Cheirogaleus medius. 

KAPPELER, P.M., OSTNER, J.: The social system of redfronted lemurs (Eulemur 

fulvus rufus) in Kirindy forest. 

KNOGGE, C.: Seed dispersal in a tamarin mixed-species troop. 

KOENIG, A., BORRIES, C., WESCHE, K.: Use of a dipterocarp forest and its consequences 

on Hanuman langurs (Presbytis entellus). 

RASOLOARISON, R.: Taxonomy and biogeographic evolution of mouse lemurs 

(Microcebus spp.). 

SCHWAB, D.: Social and mating system of the pygmy mouse lemur (Microcebus 

myoxinus). 

SCHWAB, D.: Coexistence of three sympatric lemurs (Microcebus myoxinus, Microcebus 

murinus and Cheirogaleus medius) in Kirindy forest (West Madagascar). 

ZINNER, D., TORKLER, F., PELÁEZ, F.: Habitat quality and distribution of hamadryas 

baboons in Eritrea. 

16th Ethologist Meeting, Halle (Saale), 06.-11.09.98, 

KOENIG, A., BORRIES, C.: Kosten des Gruppenlebens bei Hanuman Languren 

(Presbytis entellus). 

BORRIES, C.: Migrationsverhalten männlicher Hanuman Languren (Presbytis 

entellus). 

VII Congreso Nacional y IV Latinoamericano de Etología, Pontevedra, Peru, 07.- 

10.09.98, PELÁEZ, F., ZINNER, D., TORKLER, F.: Informe preliminar sobre la 

distribución de los papiones de desierto (Papio h. hamadryas) en Eritrea. 

IV Congreso Latinoamericano de Ecología, Arequipa, Peru, 20.-25.10.98, 

HEIDUCK, S.: Estratégia alimentar do macaco guigó (Callicebus personatus) na 

mata atlântica do Brasil. 

HEYMANN, E.W.: El rol de la presa en la diferenciación del nicho ecológico en 

pequeños platirrhinos simpátricos. 

KNOGGE, C.: Efectiva dispersión de semillas a través de los pichicos Saguinus 

mystax y Saguinus fuscicollis. 

WGL Conference: Biodiversity: Recording, Conservation, Use, Berlin, 17.-18.11.98, 

ZINNER, D.: Diversität und Erhaltung von Primaten. 

Seminar lectures 


Seminar of the Institute of Ethology and Physiology, University of Bielefeld, 

03.02.98, KNOGGE, C.: Tier-Pflanze-Interaktionen im Amazonas-Regenwald: Samenausbreitung 

durch die sympatrischen Tamarinarten Saguinus mystax und 

Saguinus fuscicollis (Callitrichinae, Primates). 





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Zoological Institute, University of Würzburg, 11.02.98, KAPPELER, P.M.: Determinanten 

der Sozialsysteme der Primaten. 

DPZ-Colloquium, Göttingen, 25.02.98, ZINNER, D.: Eritrea - ein Land für Freilandforschung 

an Primaten? 

Department of Biology, University of Asmara, Eritrea, 03.04.98, ZINNER, D.: Primate 

Diversity. 

Ethology and Socio-ecology, University of Utrecht, 21.04.98, KAPPELER, P.M.: The 

evolution of primate social systems and life histories. 

DPZ-Colloquium, Göttingen, 22.04.98, KNOGGE, C.: Faul oder fleißig? Wie sieht's 

im Garten der Zwerge aus? Über die Bedeutung von Schnurrbart- und Braunrückentamarinen 

als effektive Samenausbreiter im Amazonas-Regenwald Perus. 

DFG-Colloquium, University of Bayreuth, 19.05.98, KAPPELER, P.M.: Genetische 

Analyse der Paarungssysteme polygyner Lemuren. 

Zoological Institute, University of Würzburg, 20.05.98, KAPPELER, P.M.: Bestrafung 

im Tierreich. 

Faculty of Biology, University of Gießen, 01.07.98, HEYMANN, E.W.: Selbstmedikation 

bei Primaten. 

DPZ-Colloquium, Göttingen, 01.07.98, BORRIES, C., KOENIG, A.: Kindestötungen 

bei Hanuman Languren: Krankhaft oder eigennützig? 

Colloquium of the Institute for Social-Economy, University of Bremen, 01.08.98, 

KNOGGE C., KNOGGE, T.: Evolution und Kooperation- Analogien zwischen Ökonomie 

und Soziobiologie. 

Dept. of Biology, University of Asmara, Eritrea, 28.10.98, ZINNER, D.: Hamadryas 

Baboons of Eritrea: Distribution, Status and Ecology. 

Zoological Institute, University of Hamburg, 11.11.98, BORRIES, C., KOENIG, A.: 

Infantizid bei Hanuman Languren (Presbytis entellus): Biologisch angepaßt oder 

pathologisch. 

Facultad de Ingeniería Forestal, Universidad Nacional de la Amazonía Peruana, 

Iquitos (Peru), 17.11.98, 

HEYMANN, E.W.: Primates y ecosistemas tropicales: interacciones y dependencias. 

KNOGGE, C., TIRADO, E.R.: Dispersión de semillas por los pichicos Saguinus 

mystax y Saguinus fuscicollis. 

Zoological Institute, University of Hamburg, 25.11.98, ZINNER, D.: Paviane in 

Eritrea. Verbreitung, Ökologie und menschlicher Einfluß. 





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German-Madagasy Society, Rottenburg, 29.11.98, KAPPELER, P.M.: Lemuren Madagaskar's: 

Ursprung, Diversität, Zukunft. 

Dept. of Anthropology, SUNY at Stony Brook, USA, 04.12.98, KOENIG, A.: Socioecology 

of Hanuman langurs: Seasonal breeding, competition and fitness of folivorous, 

forest-dwelling females. 

University of Antwerp, Belgium, 21.12.98, ZINNER, D.: Baboons in Eritrea: Distribution, 

Status, Habitat. 

Other scientific activities 


• At the IIIth Kirindy-Symposium more than 50 participants exchanged results of 

and plans for research projects at our Malagasy field station. 

• E.W. Heymann co-organized a symposium on "Callitrichine mixed-species 

troops: patterns, mechanisms and consequences" at the XVIIth IPS Congress. 

• E.W. Heymann and D. Zinner are members of the IUCN/SSC Primate Specialist 

Group. 

• E.W. Heymann is a board member of the German Society for Tropical Ecology. 

• P.M. Kappeler is associate editor for Behavioral Ecology and Sociobiology and 

scientific advisor to the German Academic Exchange Service. 

• A. Koenig is a member of the board of the German Society for Primatology and editor 

of their newsletter. 

• D. Schwab is a member of the DAAD commission for Africa. 

• D. Zinner submitted a report on the situation of primates in Eritrea for the Ministry 

of Agriculture and the Department of Environment in Eritrea. 

• Members of our department prepared scientific reviews for several organizations 

(NSF, Leakey Foundation, Studienstiftung des Deutschen Volkes) and scientific 

publishers and journals (American Journal of Primatology, Animal Behaviour, Behavioral 

Ecology, Behavioral Ecology and Sociobiology, Cambridge University 

Press, Ethology, Journal of Medical Primatology, Journal of Tropical Ecology, Proceedings 

of the Royal Society: Biological Sciences, Yearbook of Physical Anthropology). 





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DEPARTMENT OF ETHOLOGY AND ECOLOGY Head of ... - DPZ

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