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184 The Trees <strong>of</strong> Great Britain and Ireland<br />
THUYA PLICATA, GIANT THUYA<br />
Tlutyaplicata, D . Don in Lambert, Pinus, ed. i, ii. 19 (1824); Masters, Card. Chron. xxi. 214;<br />
figs. 69, 70, 71 (1897); Sudworth, Check List Forest Trees U.S. 3 1 (1898); Sargent, Manual<br />
Trees JV. America, 7 5 (1905).<br />
Tlutya gigantea, Nuttall, Jour. Philad. A fad. vii. 52 (1834); Sargent, Silva N. America, x. 129, t.<br />
533 (1896); Kent, in Veitch's Man. Coniferie, 2 39 (1900).<br />
Thuya Menziesii, Douglas, ex Carriere, Traitc Gen. Conif. 1 07 (1867).<br />
Thuya Lobbi, Hort.<br />
Thuya Craigiana, Hort. \non A. Murray, Bot. Exped. Oregon, z ( 1853)].<br />
A l<strong>of</strong>ty tree, attaining a height <strong>of</strong> 200 feet, with a trunk remarkably conical, <strong>the</strong><br />
base being broad and buttressed, sometimes girthing as much as 40 to 50 feet near<br />
<strong>the</strong> ground.<br />
Bark <strong>of</strong> <strong>the</strong> trunk fissuring longitudinally in narrow thick plates, which scale <strong>of</strong>f,<br />
leaving exposed <strong>the</strong> reddish brown cortex beneath. On <strong>the</strong> branches, <strong>the</strong> bark only<br />
begins to scale when <strong>the</strong>y become old and thick. Branches horizontal, ascending<br />
towards <strong>the</strong>ir ends, forming in England a dense, narrow, pyramidal tree, usually<br />
clo<strong>the</strong>d to <strong>the</strong> base.<br />
The 3-4 pinnate branch-systems, disposed in horizontal planes, have <strong>the</strong>ir main<br />
axes terete and covered with long leaves ending in acute points which keep parallel to<br />
<strong>the</strong> axes. The glands on <strong>the</strong>se leaves are inconspicuous or absent. On <strong>the</strong> ultimate<br />
axes <strong>the</strong> leaves are smaller, <strong>the</strong> flat ones scarcely glandular, and ending in mucronate<br />
points; <strong>the</strong> lateral ones keeled on <strong>the</strong> back, slightly curved, and ending in sharp<br />
cartilaginous points. On <strong>the</strong> lower surface <strong>of</strong> most branchlets <strong>the</strong> foliage is streaked<br />
with white, some branchlets usually remaining uniformly green.<br />
The male flowers are dark red in colour, cylindrical, and composed <strong>of</strong> about<br />
6 decussate pairs <strong>of</strong> stamens.<br />
The cones when ripe do not remain erect, but are deflected out <strong>of</strong> <strong>the</strong> plane<br />
<strong>of</strong> <strong>the</strong> branchlets. They are oblong, light brown in colour, and composed <strong>of</strong> 5 to 6<br />
pairs <strong>of</strong> scales, <strong>of</strong> which <strong>the</strong> 2nd, 3rd, and 4th pairs are larger than <strong>the</strong> o<strong>the</strong>rs, and<br />
fertile. The scales are oval or spathulate, with a rounded apex, from immediately<br />
below which externally a small deltoid process is given <strong>of</strong>f. The seeds, 2 or 3 on<br />
each fertile scale, are brown in colour, two-thirds <strong>the</strong> length <strong>of</strong> <strong>the</strong> scale, and<br />
surrounded laterally by a scarious wing, which is deeply notched at its summit.<br />
Seedling. 1—The 2 cotyledons are linear, flat, acute at <strong>the</strong> apex, and slightly<br />
tapering towards <strong>the</strong> base, supported on a terete caulicle, about J inch long, which<br />
ends in a long brown flexuose primary root giving <strong>of</strong>f a few fibres. The stem, terete<br />
and smooth near <strong>the</strong> base, becomes ridged above by <strong>the</strong> decurrent leaf-bases. The<br />
first 4 true leaves are in opposite pairs, decussate with <strong>the</strong> cotyledons. Above<br />
<strong>the</strong>se <strong>the</strong> stem gives <strong>of</strong>f a number <strong>of</strong> whorls or pseudo-whorls <strong>of</strong> longer (1 inch)<br />
sharply pointed leaves, dark green above and pale beneath, with markedly decurrent<br />
1 Figured in Lubbock, Seedlings, ii. 551, hg. 676 (1892), and Sargent, lee. cil. t. 533, fig. 1 2.<br />
Thuyi<br />
bases. After a few <strong>of</strong> <strong>the</strong>se whorls lateral branches are given <strong>of</strong>f, which sometimes<br />
bear a few acicular leaves at <strong>the</strong>ir bases. The lateral branches ramify and approach<br />
in character those <strong>of</strong> <strong>the</strong> adult plant, as <strong>the</strong> leaves are arranged decussately in 4<br />
ranks. These leaves are variable, being acicular and loosely imbricated, or scale-like<br />
and closely imbricate. The branches are ascending, horizontal, or drooping, and are<br />
more or less flattened from above downwards.<br />
HISTORY 1<br />
This tree was discovered by Nee, who accompanied Malaspina in his voyage round<br />
<strong>the</strong> world during <strong>the</strong> years 1789 to 1794; and his specimen, ga<strong>the</strong>red at Nootka<br />
Sound, is preserved in <strong>the</strong> Natural History Museum at South Kensington. It was<br />
referred to by James Donn, in Hortus Cantab, ed. 4 (1807), as Thuya plicata, without<br />
any description; and subsequently D. Don drew up from it <strong>the</strong> oldest description <strong>of</strong><br />
<strong>the</strong> species under <strong>the</strong> same name. The Thuya plicata <strong>of</strong> gardens, which was early<br />
in cultivation, is a variety <strong>of</strong> Thuya occidentalis, and has no connection with <strong>the</strong><br />
plant <strong>of</strong> Nee.<br />
Archibald Menzies, who accompanied Vancouver's expedition as botanist,<br />
ga<strong>the</strong>red specimens also at Nootka Sound in 1795. Nuttall received specimens<br />
later from <strong>the</strong> Fla<strong>the</strong>ad river, on which he founded his description <strong>of</strong> <strong>the</strong> species as<br />
Thuya gigantea. It was introduced into cultivation 2 in 1853 by W. Lobb, and<br />
distributed from Veitch's nursery at Exeter as Thuya Lobbi, as at that time Nuttall's<br />
name Thuya gigantea was wrongly applied to Libocedrus decurrens, and Don's name,<br />
Thuya plicata, in a similar erroneous way, had gone into common use for a variety<br />
<strong>of</strong> Thuya occidentalis. Afterwards <strong>the</strong> tree became generally known in England as<br />
Thuya gigantea; and it is unfortunate that Don's name, Thuya plicata, must,<br />
following <strong>the</strong> law <strong>of</strong> priority, be substituted for a name so well known and so<br />
established as gigantea. This change <strong>of</strong> name has, however, been adopted in <strong>the</strong><br />
Kew Hand List <strong>of</strong> Conifers, and by Sudworth and Sargent in North America,<br />
and on <strong>the</strong> whole it is now most convenient to adopt <strong>the</strong> name Thuya plicata.<br />
(A. H.)<br />
DISTRIBUTION<br />
This tree is, next to <strong>the</strong> Douglas fir, <strong>the</strong> most important from an economic point<br />
<strong>of</strong> view in nor<strong>the</strong>rn Oregon, Washington, and British Columbia.<br />
It extends in <strong>the</strong> north as far as sou<strong>the</strong>rn Alaska, in <strong>the</strong> east to <strong>the</strong> Cceur<br />
d'Alene Mountains in Idaho and to north-western Montana, and in <strong>the</strong> south to<br />
Mendocino County in nor<strong>the</strong>rn California. It is known as Cedar, or Red Cedar, and<br />
is found most abundantly on wet soils and in wet climates, ascending from sea level<br />
to an elevation, according to Sargent, <strong>of</strong> 6000 feet, where it becomes a low shrub.<br />
1 See Masters, in Card. Chron. Inc. cit.<br />
2 At <strong>the</strong> Royal Botanic Gardens, Edinburgh, <strong>the</strong>re were in 1884 five <strong>trees</strong> <strong>of</strong> supposed Thuya gigantea, which were<br />
raised, it was said, from seed sent to Edinburgh by Jeffrey in 1851, while collecting for <strong>the</strong> Oregon Association. Three <strong>of</strong><br />
<strong>the</strong>se <strong>trees</strong>, according to Nicholson, were true giganlea, <strong>the</strong> o<strong>the</strong>r two being what is now known as Thuya occidentalis, var.<br />
plicata. See Woods and forests, Feb. 27, and Mar. 19, 1884. These <strong>trees</strong> cannot now be identified.<br />
I 2 B