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late pleistocene population interaction in western europe

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may be used to test Kle<strong>in</strong>’s (1989) proposition that postdepositional breakage would be<br />

more prevalent, <strong>in</strong> general, on large versus small ungu<strong>late</strong>s.<br />

Controll<strong>in</strong>g for spatial variation as much as possible, the <strong>in</strong>cidence of cracks on<br />

long bones <strong>in</strong> the best-preserved rows of Sa<strong>in</strong>t-Césaire is much greater for horse and<br />

bison than for re<strong>in</strong>deer (Table 42). These differences with re<strong>in</strong>deer are significant (horse:<br />

ts = 6.75, p < 0.0001; bison: ts = 6.14, p < 0.0001, mean of meter 2 and 3), but not<br />

between horse and bison (ts = 0.83, p < 0.41). These data imply that horse and bison<br />

bones would be more likely to crack and break, possibly as a result of scal<strong>in</strong>g effects, and<br />

therefore more difficult to identify, than those of re<strong>in</strong>deer. One such effect might be<br />

associated with bone thickness and differential moisture. Because the gradient of<br />

moisture appears to be greater <strong>in</strong> large animal bones than <strong>in</strong> smaller ones (D. Fisher, pers.<br />

com. 2003), fiber contraction and dilatation might be <strong>in</strong>creased <strong>in</strong> the bone walls of the<br />

former taxa, which would promote crack<strong>in</strong>g.<br />

Whatever is the cause of this <strong>in</strong>creased fragmentation, these results suggest that<br />

large ungu<strong>late</strong> limbs may be under-identified relative to those of re<strong>in</strong>deer at Sa<strong>in</strong>t-<br />

Césaire. This f<strong>in</strong>d<strong>in</strong>g has another important implication. Because teeth are highly<br />

diagnostic, this part is more likely to be over-represented with respect to limbs <strong>in</strong> large<br />

ungu<strong>late</strong>s than <strong>in</strong> smaller taxa. This might expla<strong>in</strong> why head parts are so abundant <strong>in</strong><br />

some assemblages dom<strong>in</strong>ated by large ungu<strong>late</strong>s (e.g., Kle<strong>in</strong> 1989).<br />

223

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