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Bottom Trawl Surveys - Proceedings of a Workshop Held at Ottawa ...

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114<br />

It is seldom the case th<strong>at</strong> survey results<br />

are used on a "stand alone" basis; more <strong>of</strong>ten<br />

they are used as independent corrobor<strong>at</strong>ion <strong>of</strong><br />

results obtained through other methods, such as<br />

tagging, c<strong>at</strong>ch-curve analyses or various classes<br />

<strong>of</strong> yield models. If the survey design fits the<br />

characteristics <strong>of</strong> the study organism and the<br />

d<strong>at</strong>a are analyzed appropri<strong>at</strong>ely, then the<br />

results should valid<strong>at</strong>e trends and parameter<br />

estim<strong>at</strong>es derived elsewhere, or they should<br />

provide some altern<strong>at</strong>e explan<strong>at</strong>ion consistent<br />

with stock <strong>at</strong>tributes. Where designs have only<br />

minor deficiencies resulting from peculiarities<br />

<strong>of</strong> sampling situ<strong>at</strong>ions, the investig<strong>at</strong>or should<br />

provide some qualit<strong>at</strong>ive or quantit<strong>at</strong>ive<br />

assessment <strong>of</strong> the bias which may result.<br />

DESIGN OF THE SURVEY<br />

Many <strong>of</strong> the criticisms directed <strong>at</strong> surveys<br />

as they are normally practised cou I d be<br />

mitig<strong>at</strong>ed if sufficient time was expended<br />

beforehand in precisely defining the objectives<br />

<strong>of</strong> the survey and reviewing relevant inform<strong>at</strong>ion<br />

on problems specific to the study. Failure to<br />

do so <strong>of</strong>ten results in surveys which are <strong>of</strong><br />

little direct use to the resource scientist<br />

because confidence limits are unacceptably<br />

large, etc. Such surveys are then justified on<br />

the basis th<strong>at</strong> they may be "good indic<strong>at</strong>ors <strong>of</strong><br />

annual trends", whereas wh<strong>at</strong> the scientist<br />

really needs is a good estim<strong>at</strong>e <strong>of</strong> the absolute<br />

value <strong>of</strong> the parameter being measured. Failure<br />

to produce such estim<strong>at</strong>es can lead to articles<br />

in the industry press, such as those in N<strong>at</strong>ional<br />

Fisherman (June 1979).<br />

Pre-survey activities should include<br />

formul<strong>at</strong>ing a clear st<strong>at</strong>ement <strong>of</strong> the objectives;<br />

i.e., is the survey to measure abundance,<br />

determine size or age composition, deline<strong>at</strong>e<br />

distribution <strong>of</strong> various segments <strong>of</strong> the stock,<br />

determine the diel activities <strong>of</strong> the species, or<br />

determine some combin<strong>at</strong>ion <strong>of</strong> these and other<br />

elements? This process should help the scien­<br />

tist assign the level <strong>of</strong> sampling wh·ich may be<br />

necessary <strong>at</strong> each st<strong>at</strong>ion. For instance, sampling<br />

intensity for stock segreg<strong>at</strong>ion studies<br />

would be considerably less than th<strong>at</strong> required<br />

for the determin<strong>at</strong>ion <strong>of</strong> diel behaviour<br />

p<strong>at</strong>terns.<br />

The other major pre-survey activity should<br />

be a coll<strong>at</strong>ion <strong>of</strong> available inform<strong>at</strong>ion which<br />

may be relevant to the variable under study.<br />

For example, any known characteristics <strong>of</strong><br />

distribution or environment may be critical<br />

determinants <strong>of</strong> the eventual survey design and<br />

should be reviewed, even if only dealing <strong>at</strong> the<br />

generic or ordinal level <strong>of</strong> the subject species.<br />

A gre<strong>at</strong> deal <strong>of</strong> time and effort in the field may<br />

be saved through this process not to mention the<br />

obvious impact on results.<br />

Two <strong>of</strong> the primary constraints facing the<br />

resource scientist are the time or funding<br />

available with which to conduct the survey.<br />

Because funding is generally limited, the<br />

scientist may decide to collect the maximum<br />

possible amount <strong>of</strong> inform<strong>at</strong>ion, hoping th<strong>at</strong> some<br />

compens<strong>at</strong>ion between quality and quantity <strong>of</strong><br />

d<strong>at</strong>a will occur. In other words, the<br />

investig<strong>at</strong>or may consider th<strong>at</strong> he has little<br />

time available in which to modify the survey<br />

based on real-time results, particularly if the<br />

resource/area eel I has received little previous<br />

<strong>at</strong>tention. This is generally the case for widely<br />

distributed resources; thus, the investig<strong>at</strong>or<br />

may be unable to quickly sample a<br />

"represent<strong>at</strong>ive" segment <strong>of</strong> the range. When<br />

this is true, caution must be exercised to avoid<br />

interpreting local fe<strong>at</strong>ures as general<br />

popul<strong>at</strong>ion characteristics. There may also be<br />

prob I ems due to i nterspecitic differences in<br />

dispersion or behavioural p<strong>at</strong>terns and<br />

intraspecitic seasonal eftects, such as<br />

migr<strong>at</strong>ions or spawning aggreg<strong>at</strong>ions.<br />

Unfortun<strong>at</strong>ely, the investig<strong>at</strong>or seldom has<br />

sufficient resources to deline<strong>at</strong>e these temporal<br />

aspects, particularly for widespread stocks.<br />

For most, it' not all, groundfish stocks,<br />

there is some degree <strong>of</strong> overdispersion within<br />

the popul<strong>at</strong>ion; i.e., the distribution <strong>of</strong><br />

individuals is contagious r<strong>at</strong>her than regular or<br />

random. It may be confined to a grouping <strong>of</strong><br />

popul<strong>at</strong>ion sub-units in response to<br />

environmental fe<strong>at</strong>ures, such as substr<strong>at</strong>e<br />

availability or w<strong>at</strong>er mass <strong>at</strong>tributes, or there<br />

may be distinct groupings within any homogeneous<br />

area. In the former case, there may be random<br />

distribution <strong>of</strong> sub-units within which there is<br />

regular or random dispersion, while in the<br />

l<strong>at</strong>ter there will be contagious distribution <strong>of</strong><br />

individuals. Contagious distributions may thus<br />

arise in a number <strong>of</strong> ways and the size <strong>of</strong><br />

sampling units must be appropri<strong>at</strong>e not only to<br />

total area coverage but also to the magnitude <strong>of</strong><br />

the sp<strong>at</strong>ial dispersion. The effects <strong>of</strong> the size<br />

<strong>of</strong> the sampling unit rel<strong>at</strong>ive to heterogeneity<br />

in distribution will be discussed l<strong>at</strong>er.<br />

It is uncommon th<strong>at</strong> the investig<strong>at</strong>or will<br />

be totally ignorant <strong>of</strong> the basic distributional<br />

properties <strong>of</strong> the resource. It is therefore<br />

possible to postul<strong>at</strong>e a distribution and design<br />

the survey not only to collect desired<br />

biological inform<strong>at</strong>ion but also to test the<br />

di stributi anal hypothesis. In postul<strong>at</strong>ing the<br />

distribution, the investig<strong>at</strong>or should be<br />

prepared to justify the assumptions made. For<br />

example, if a species <strong>of</strong> fl<strong>at</strong>fish is thought to<br />

be contagiously distributed in rel<strong>at</strong>ion to the<br />

presence <strong>of</strong> mud bottom, the hypothetical<br />

probability distribution must also gener<strong>at</strong>e<br />

frequencies <strong>of</strong> occurrence which reflect this<br />

substr<strong>at</strong>e distribution.<br />

Having postul<strong>at</strong>ed the distribution, the<br />

investig<strong>at</strong>or must then design the survey<br />

appropri<strong>at</strong>e to it. It is <strong>at</strong> this stage th<strong>at</strong> the<br />

aforementioned fiscal and temporal constraints<br />

generally manifest themselves. The basic<br />

concern <strong>of</strong> the scientist will be either to<br />

obtain maximum precision <strong>of</strong> an estim<strong>at</strong>e (e.g.,<br />

popul<strong>at</strong>ion mean) within a certain budget or to<br />

minimize the cost (generally with regard to<br />

areal coverage) for a specified precision. In<br />

two-stage sampling where the selection <strong>of</strong>

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