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Bottom Trawl Surveys - Proceedings of a Workshop Held at Ottawa ...

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138<br />

In the area south <strong>of</strong> 5o·N, the same series<br />

<strong>of</strong> biomass estim<strong>at</strong>es was made except th<strong>at</strong> with<br />

the limited number <strong>of</strong> str<strong>at</strong>a the combin<strong>at</strong>ion <strong>of</strong><br />

small str<strong>at</strong>a into depth levels was not used<br />

(Table 8).<br />

In considering the whole area <strong>of</strong> the<br />

Esquiman Channel, the same series <strong>of</strong> estim<strong>at</strong>es<br />

was examined in the northern area except th<strong>at</strong><br />

with the two larger areas a further estim<strong>at</strong>e was<br />

available by considering the whole area as made<br />

up <strong>of</strong> two str<strong>at</strong>a (Table 6).<br />

Interpret<strong>at</strong>ion <strong>of</strong> the results is<br />

complic<strong>at</strong>ed by the fact th<strong>at</strong> different str<strong>at</strong>um<br />

areas were used between the old and the new<br />

str<strong>at</strong>a, and this affects both the estim<strong>at</strong>es <strong>of</strong><br />

the mean/unit area and the expansion <strong>of</strong> this<br />

mean to biomass. However, within the various<br />

combin<strong>at</strong>ions <strong>of</strong> the small str<strong>at</strong>a a trend seems<br />

to be clear in the northern and total area,<br />

namely, th<strong>at</strong> with the increased number <strong>of</strong> str<strong>at</strong>a<br />

the lower mean is obtained and this is<br />

accompanied by an increase in precision as<br />

indic<strong>at</strong>ed both by the variance and by the<br />

confidence limits assigned to the biomass<br />

estim<strong>at</strong>es after areal expansion. In the<br />

southern part <strong>of</strong> the area, the str<strong>at</strong>ified<br />

sampling was poor, with only three out <strong>of</strong> the<br />

seven str<strong>at</strong>a sampled enough to yield a variance<br />

estim<strong>at</strong>e and two str<strong>at</strong>a not sampled <strong>at</strong> all.<br />

Thus, it is not surprising th<strong>at</strong> the best<br />

precision was obtained by considering the area<br />

as a single homogeneous one and sampling <strong>at</strong><br />

random within it.<br />

With regard to the different estim<strong>at</strong>es and<br />

the possible bias th<strong>at</strong> may exist in each, we are<br />

in a quandary, as in each method <strong>of</strong> computing<br />

the mean we have used different assumptions.<br />

Intuitively, in this instance where we are<br />

using a fairly large number <strong>of</strong> small str<strong>at</strong>a<br />

which we would expect to be reasonably<br />

homogeneous, we might expect th<strong>at</strong> the "best"<br />

estim<strong>at</strong>e might be obtained from using the<br />

gre<strong>at</strong>est number <strong>of</strong> str<strong>at</strong>a; however, as we might<br />

also expect from the lack <strong>of</strong> differences between<br />

str<strong>at</strong>um means within the same depth zone, the<br />

difference in estim<strong>at</strong>es between this one and the<br />

estim<strong>at</strong>e derived from the str<strong>at</strong>a combined by<br />

depth is trivial. In this case, we think we<br />

could conclude th<strong>at</strong> the "best estim<strong>at</strong>e" <strong>of</strong><br />

biomass for the total area might be 12,000 ±<br />

2000 metric tons in 1975.<br />

We can also conclude th<strong>at</strong> combining str<strong>at</strong>a<br />

within depth zones is acceptable though clearly<br />

when the larger str<strong>at</strong>a <strong>of</strong> the new system <strong>of</strong><br />

str<strong>at</strong>ific<strong>at</strong>ion are used there is some widening<br />

<strong>of</strong> the 95% confidence interval.<br />

A comparison between the estim<strong>at</strong>es <strong>of</strong><br />

mean/unit area obtained in different years is<br />

also instructive (Table g), This table displays<br />

the results obtained in each str<strong>at</strong>um when the<br />

new str<strong>at</strong>ific<strong>at</strong>ion method was uniformly applied<br />

through the series; thus, vari<strong>at</strong>ion due to the<br />

use <strong>of</strong> different areal weighting is removed, and<br />

the estim<strong>at</strong>es therefore represent sampling<br />

variability and changes in biomass between<br />

years. An examin<strong>at</strong>ion <strong>of</strong> the means suggests<br />

th<strong>at</strong> we should not put too much emphasis on<br />

sampling the deepest str<strong>at</strong>um (115) because <strong>of</strong><br />

the low shrimp abundance and the rel<strong>at</strong>ively<br />

small area involved, while a perusal <strong>of</strong> the<br />

coefficients <strong>of</strong> vari<strong>at</strong>ion suggests th<strong>at</strong> sampling<br />

in Str<strong>at</strong>a 113 and 114 is likely not too bad and<br />

th<strong>at</strong> any improvements are most likely<br />

forthcoming from improving the sampling in<br />

Str<strong>at</strong>um 112. From a knowledge <strong>of</strong> the<br />

distribution <strong>of</strong> shrimp, it is likely th<strong>at</strong> an<br />

improvement would be obtained by refining the<br />

depth str<strong>at</strong>ific<strong>at</strong>ion used and alloc<strong>at</strong>ing sets in<br />

proportion to observed variability in each<br />

str<strong>at</strong>um.<br />

PROBLEMS ASSOCIATED WITH DIURNAL VARIABILITY<br />

In the biomass surveys in the Northeast<br />

Gulf <strong>of</strong> St. Lawrence, sampling has been<br />

restricted to the daylight hours; however, in<br />

the <strong>of</strong>fshore areas <strong>of</strong>f Labrador and in the Davis<br />

Strait area this has not been the case.<br />

The vertical migr<strong>at</strong>ion <strong>of</strong> shrimp has been<br />

an accepted fact for many years. At various<br />

times during the 24-hour cycle, especially <strong>at</strong><br />

night, shrimp move <strong>of</strong>f the bottom and disperse<br />

to considerable distances in the w<strong>at</strong>er column.<br />

Sampling the stocks with "bottom" trawls other·<br />

than <strong>at</strong> times <strong>of</strong> optimum density should lead to<br />

an underestim<strong>at</strong>ion <strong>of</strong> stock size since those<br />

animals occurring outside the range <strong>of</strong> the<br />

trawl's vertical opening will not be accounted<br />

for. Daylight sampling alone tends to be<br />

inefficient, mainly because the migr<strong>at</strong>ion <strong>of</strong>ten<br />

extends well into the daylight hours and,<br />

therefore, the problem is not resolved.<br />

Intuitively, any adjustment for diel<br />

variability should have <strong>at</strong> least two reasonably<br />

pr<strong>of</strong>ound effects: (a) the biomass should<br />

increase; and (b) the variance associ<strong>at</strong>ed with<br />

the mean biomass should be reduced.<br />

Attempts to achieve these effects have been<br />

made but the d<strong>at</strong>a base is <strong>of</strong>ten rudimentary.<br />

C<strong>at</strong>ch d<strong>at</strong>a from a survey in the Davis Strait in<br />

1979 (Parsons 1979) were adjusted to account for<br />

diel variability: (a) observed from 24-hour<br />

sampling during the survey (Figure 10); and (b)<br />

according to p<strong>at</strong>terns observed for the month <strong>of</strong><br />

September from more extensive commercial d<strong>at</strong>a<br />

(Carlsson et al. 1978). In both cases, the<br />

desired results were achieved (Table 10) with<br />

higher biomass and finer confidence intervals<br />

associ<strong>at</strong>ed with the p<strong>at</strong>tern observed <strong>at</strong> the time<br />

<strong>of</strong> the survey. The exercise, however, did not<br />

improve the d<strong>at</strong>a enough to give values <strong>of</strong> the<br />

lower limits (95%) gre<strong>at</strong>er than zero.

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