A Criticism of the Cell-Theory; being an Answer to Mr. Sedgwick's ...
A Criticism of the Cell-Theory; being an Answer to Mr. Sedgwick's ...
A Criticism of the Cell-Theory; being an Answer to Mr. Sedgwick's ...
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A CRITICISM OP THE CELL-THEORY. 155<br />
Although in this passage, which is descriptive <strong>of</strong> Thallophytes,<br />
Goebel attaches <strong>to</strong>o much import<strong>an</strong>ce, as I think, <strong>to</strong> <strong>the</strong> continuity<br />
<strong>of</strong> a vesicle as determining <strong>the</strong> unicellularity <strong>of</strong> a pl<strong>an</strong>t,<br />
he shows clearly enough that he regards <strong>the</strong> growth <strong>an</strong>d mode<br />
<strong>of</strong> extension <strong>of</strong> <strong>the</strong> pro<strong>to</strong>plasm, not its division in<strong>to</strong> cells, as<br />
<strong>the</strong> feature <strong>of</strong> fundamental import<strong>an</strong>ce.<br />
There is <strong>the</strong> fur<strong>the</strong>r property in pl<strong>an</strong>ts that continuity<br />
between <strong>the</strong> cells <strong>of</strong> highly org<strong>an</strong>ised multicellular pl<strong>an</strong>ts has<br />
been shown <strong>to</strong> be <strong>of</strong> very general, if not universal, occurrence.<br />
And if complete separation were <strong>to</strong> be insisted upon as a<br />
characteristic <strong>of</strong> a cell, <strong>an</strong>y given Angiosperm, or o<strong>the</strong>r highly<br />
org<strong>an</strong>ised pl<strong>an</strong>t, could no longer be considered as <strong>an</strong> aggregate<br />
<strong>of</strong> life units, but ra<strong>the</strong>r as a conjunct mass <strong>of</strong> pro<strong>to</strong>plasm,<br />
imperfectly broken up in<strong>to</strong> corpuscles, in each <strong>of</strong> which <strong>the</strong>re<br />
is a nucleus. It is but a step from <strong>the</strong> much-br<strong>an</strong>ched, multinucleate<br />
Coeloblastse, which have no partitions, <strong>to</strong> <strong>the</strong> forma,<br />
tion <strong>of</strong> incomplete partitions, breaking up <strong>the</strong> pro<strong>to</strong>plasm in<strong>to</strong><br />
small masses, which remain, however, linked with one <strong>an</strong>o<strong>the</strong>r,<br />
<strong>an</strong>d so preserve <strong>an</strong> original continuity similar <strong>to</strong> that <strong>of</strong> <strong>the</strong><br />
Coeloblastse, which has only apparently but never actually been<br />
broken.<br />
So much has this idea impressed itself on <strong>the</strong> minds <strong>of</strong> some<br />
observers, that H<strong>of</strong>meister suggested that <strong>the</strong> creeping motion<br />
<strong>of</strong> <strong>the</strong> plasmodia <strong>of</strong> <strong>the</strong> Myxomycetes <strong>an</strong>d <strong>the</strong>ir later tr<strong>an</strong>sfer,<br />
mation in<strong>to</strong> fructification, is representative <strong>of</strong> <strong>the</strong> simplest type<br />
<strong>of</strong> growth, even for more highly org<strong>an</strong>ised pl<strong>an</strong>ts. This<br />
opinion has been quoted with approval by von Sachs, who,<br />
before even <strong>the</strong> continuity <strong>of</strong> <strong>the</strong> pro<strong>to</strong>plasm <strong>of</strong> pl<strong>an</strong>t cells<br />
was established, wrote that " fundamentally every pl<strong>an</strong>t, how-*<br />
ever highly org<strong>an</strong>ised, is a pro<strong>to</strong>plasmic body, coherent in<br />
itself, which, clo<strong>the</strong>d without by a cell-wall <strong>an</strong>d traversed<br />
internally by innumerable partitions, grows; <strong>an</strong>d it appears<br />
that <strong>the</strong> more vigorously this formation <strong>of</strong> chambers <strong>an</strong>d<br />
walls proceeds with <strong>the</strong> nutrition <strong>of</strong> <strong>the</strong> pro<strong>to</strong>plasm, <strong>the</strong> higher<br />
also is <strong>the</strong> development attained by <strong>the</strong> <strong>to</strong>tal org<strong>an</strong>isation."<br />
Expressed in this way, <strong>the</strong> phenomenon <strong>of</strong> cell-formation is<br />
represented <strong>to</strong> us as <strong>being</strong> nothing more th<strong>an</strong> a particular