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A Criticism of the Cell-Theory; being an Answer to Mr. Sedgwick's ...

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A CRITICISM OF THE CELL-THEORY. 173<br />

from experience, for not long since I was much puzzled by<br />

such a reticulum, <strong>an</strong>d had I been less cautious I should have<br />

published, as a great morphological discovery, statements which<br />

rested on a wholly insufficient basis <strong>of</strong> experience. The subject<br />

requires fur<strong>the</strong>r investigation, <strong>an</strong>d <strong>the</strong> most that one c<strong>an</strong> say<br />

now is, that it is possible that <strong>Mr</strong>. Sedgwick, good observer as<br />

he is, may have been mistaken. And he will pardon my<br />

observing that <strong>the</strong> things which he states are not " facts."<br />

They are his own inferences from his own individual observations,<br />

<strong>an</strong>d will require very abund<strong>an</strong>t confirmation before <strong>the</strong>y<br />

c<strong>an</strong> take r<strong>an</strong>k as what we agree <strong>to</strong> regard as " facts." All <strong>the</strong><br />

" facts " we have at present, i. e. <strong>the</strong> accumulated observations<br />

<strong>of</strong> hundreds <strong>of</strong> highly-trained <strong>an</strong>d able observers, are fundamentally<br />

opposed <strong>to</strong> <strong>an</strong>y such account <strong>of</strong> pro<strong>to</strong>plasmic growth<br />

apart from nuclear formation as <strong>Mr</strong>. Sedgwick gives us. But<br />

<strong>the</strong>re is <strong>an</strong>o<strong>the</strong>r way <strong>of</strong> looking at it, namely, that he has only<br />

overstated his case, <strong>an</strong>d that <strong>the</strong> growth <strong>of</strong> <strong>the</strong> tissues in<br />

question resembles <strong>the</strong> apparent creeping motion <strong>of</strong> <strong>the</strong> plasmodia<br />

<strong>of</strong> <strong>the</strong> Myxomycetes. That this may be <strong>the</strong> case is<br />

supported by a study <strong>of</strong> <strong>Mr</strong>. Asshe<strong>to</strong>n's recent account <strong>of</strong> <strong>the</strong><br />

growth <strong>of</strong> <strong>the</strong> mesoblast <strong>an</strong>d <strong>of</strong> <strong>the</strong> inner layer <strong>of</strong> <strong>the</strong> epiblast<br />

in <strong>the</strong> embryo <strong>of</strong> <strong>the</strong> rabbit. It presents no <strong>the</strong>oretical difficulties,<br />

but it should be remarked that <strong>Mr</strong>. Asshe<strong>to</strong>n figures<br />

numerous nuclei at <strong>the</strong> very edge <strong>of</strong> <strong>the</strong> growing part <strong>of</strong> his<br />

reticula, which is conson<strong>an</strong>t with what we know <strong>of</strong> pro<strong>to</strong>plasmic<br />

growth in o<strong>the</strong>r cases, but not with <strong>Mr</strong>. <strong>Sedgwick's</strong><br />

account.<br />

But if <strong>Mr</strong>. Sedgwick c<strong>an</strong> prove that <strong>the</strong> reticulum is <strong>the</strong>re<br />

<strong>an</strong>d that it grows <strong>an</strong>d spreads far from <strong>the</strong> nuclei which subsequently<br />

migrate in<strong>to</strong> it, he must not suppose, as he is<br />

apparently so ready <strong>to</strong> assume, that <strong>the</strong> inveterate prejudice<br />

<strong>of</strong> morphologists will prevent <strong>the</strong>ir accepting his conclusions<br />

because <strong>of</strong> <strong>the</strong>ir <strong>the</strong>oretical difficulties. If his case is proved,<br />

it will be accepted, but he must prove it up <strong>to</strong> <strong>the</strong> hilt.<br />

And if he does prove it, what <strong>the</strong>n ? It will be <strong>an</strong> isolated<br />

case, <strong>of</strong> secondary signific<strong>an</strong>ce: merely <strong>an</strong>o<strong>the</strong>r addition <strong>to</strong><br />

our experience <strong>of</strong> <strong>the</strong> very various phenomena displayed in

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