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A Criticism of the Cell-Theory; being an Answer to Mr. Sedgwick's ...

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A CRITICISM OF THE CELL-THEORY. 169<br />

As a physiological concept it is hardly less useful, though<br />

reflection may induce us <strong>to</strong> ab<strong>an</strong>don <strong>the</strong> " cell-republic"<br />

<strong>the</strong>ory, as, indeed, it has been tacitly ab<strong>an</strong>doned by m<strong>an</strong>y.<br />

I take it that <strong>the</strong> scheme <strong>of</strong> von Sachs very nearly expresses,<br />

in general terms, <strong>the</strong> physiological import<strong>an</strong>ce <strong>of</strong> <strong>the</strong> cell.<br />

An org<strong>an</strong>ism is a pro<strong>to</strong>plasmic body, coherent in itself, which<br />

grows, <strong>an</strong>d as it grows it is divided by cleavage in<strong>to</strong> innumerable<br />

corpuscles, <strong>an</strong>d it appears that <strong>the</strong> more vigorously<br />

this formati<strong>an</strong> <strong>of</strong> corpuscles proceeds with <strong>the</strong> nutrition <strong>of</strong> <strong>the</strong><br />

org<strong>an</strong>ism, <strong>the</strong> higher also is <strong>the</strong> development attained by <strong>the</strong><br />

<strong>to</strong>tal org<strong>an</strong>isation. Nor does this statement st<strong>an</strong>d in <strong>an</strong>y contradiction<br />

<strong>to</strong> <strong>the</strong> original <strong>the</strong>ory <strong>of</strong> Schw<strong>an</strong>n, from whom I<br />

may quote two more passages : "The elementary parts <strong>of</strong> all<br />

tissues are formed <strong>of</strong> cells, in <strong>an</strong> <strong>an</strong>alogous though very<br />

diversified m<strong>an</strong>ner, so that it may be asserted that <strong>the</strong>re is<br />

one universal principle <strong>of</strong> development for <strong>the</strong> elementary<br />

parts <strong>of</strong> org<strong>an</strong>isms, however different, <strong>an</strong>d that this principle<br />

is <strong>the</strong> formation <strong>of</strong> cells." And again, he says <strong>of</strong> <strong>the</strong> relations<br />

<strong>of</strong> cells <strong>to</strong> one <strong>an</strong>o<strong>the</strong>r, " Each cell is within certain limits <strong>an</strong><br />

individual, <strong>an</strong> independent whole. The vital phenomena <strong>of</strong><br />

one are repeated, entirely or in part, in all <strong>the</strong> rest. These<br />

individuals, however, are not r<strong>an</strong>ged side by side as a mere<br />

aggregate, but so. operate <strong>to</strong>ge<strong>the</strong>r in a m<strong>an</strong>ner unknown <strong>to</strong><br />

us, as <strong>to</strong> produce a harmonious whole." It should be remembered<br />

that Schw<strong>an</strong>n regarded cells as so m<strong>an</strong>y separate vesicles,<br />

<strong>an</strong>d when allow<strong>an</strong>ce is made for this error, <strong>the</strong> second<br />

part <strong>of</strong> <strong>the</strong> last passage must be allowed <strong>to</strong> have great signific<strong>an</strong>ce.<br />

The subordination <strong>of</strong> <strong>the</strong> parts <strong>to</strong> <strong>the</strong> harmonious<br />

whole, leading <strong>to</strong> <strong>the</strong> loss <strong>of</strong> individuality <strong>of</strong> <strong>the</strong> parts, in<br />

<strong>an</strong>imal tissues, was insisted on by Hackel in his ' Generelle<br />

Morphologic' The first <strong>of</strong> <strong>the</strong> two sentences which I have<br />

quoted from Schw<strong>an</strong>n is even more true <strong>to</strong>-day th<strong>an</strong> when it<br />

was written, for we have got rid <strong>of</strong> <strong>the</strong> cell-forming matrix,<br />

<strong>the</strong> cy<strong>to</strong>blastema ; <strong>an</strong>d I would wish <strong>to</strong> insist on this passage<br />

as expressing in <strong>the</strong> clearest possible l<strong>an</strong>guage <strong>the</strong> cell-<strong>the</strong>ory<br />

as we underst<strong>an</strong>d it <strong>to</strong>-day.<br />

From this st<strong>an</strong>dpoint we c<strong>an</strong> see, obscurely it may be, why

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