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A Criticism of the Cell-Theory; being an Answer to Mr. Sedgwick's ...

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A CRITICISM OP THE DELL-THEORY. 139<br />

It is probably a fair summary <strong>of</strong> his position <strong>to</strong> say that, for<br />

<strong>the</strong> present, he limits his objections <strong>to</strong> <strong>the</strong> application <strong>of</strong> <strong>the</strong><br />

cell-<strong>the</strong>ory <strong>to</strong> <strong>the</strong> process <strong>of</strong> growth during embryonic development<br />

j but that he scarcely conceals his preference for <strong>the</strong><br />

view that <strong>the</strong>re are no such things as discrete cells in <strong>the</strong><br />

so-called multicellular org<strong>an</strong>ism. And as it is necessary, at<br />

<strong>the</strong> outset, <strong>to</strong> have a perfectly clear idea <strong>of</strong> his me<strong>an</strong>ing, I will<br />

quote passages from <strong>the</strong> work <strong>to</strong> which he refers in his opening<br />

paragraph, assuming that what he stated <strong>the</strong>n he is prepared<br />

<strong>to</strong> adhere <strong>to</strong> now, <strong>an</strong>d that his last article is intended <strong>to</strong><br />

emphasise <strong>the</strong> views which he formerly propounded, <strong>an</strong>d <strong>to</strong><br />

bring fresh evidence in support <strong>of</strong> <strong>the</strong>m.<br />

On p. 204 <strong>of</strong> <strong>the</strong> second part <strong>of</strong> his account <strong>of</strong> <strong>the</strong> development<br />

<strong>of</strong> <strong>the</strong> Cape species <strong>of</strong> Peripatus, he says:—"It is<br />

becoming more <strong>an</strong>d more clear every day that <strong>the</strong> cells composing<br />

<strong>an</strong>imal tissues are not isolated units, but that <strong>the</strong>y are<br />

connected with one <strong>an</strong>o<strong>the</strong>r. I need only refer <strong>to</strong> <strong>the</strong> connection<br />

known <strong>to</strong> exist between connective tissue cells, cartilage<br />

cells, epi<strong>the</strong>lial cells, &c. And not only may <strong>the</strong> cells <strong>of</strong> one<br />

tissue be continuous with one <strong>an</strong>o<strong>the</strong>r, but <strong>the</strong>y may also be<br />

continuous with <strong>the</strong> cells <strong>of</strong> o<strong>the</strong>r tissues. ... It is true<br />

that <strong>the</strong> cells <strong>of</strong> blood <strong>an</strong>d lymph <strong>an</strong>d <strong>the</strong> ripe generative cells<br />

are completely isolated. But <strong>the</strong> former, in <strong>the</strong>ir first stages<br />

<strong>of</strong> growth, form part <strong>of</strong> <strong>the</strong> syncytium, as in all probability do<br />

<strong>the</strong> latter also. This continuity, which for a priori reasons<br />

we should expect, has hi<strong>the</strong>r<strong>to</strong> been regarded as a fact <strong>of</strong> little<br />

morphological import<strong>an</strong>ce <strong>an</strong>d relegated <strong>to</strong> <strong>the</strong> category <strong>of</strong><br />

secondary features. The ovum, it is said, segments in<strong>to</strong><br />

completely isolated cells, <strong>an</strong>d <strong>the</strong> connection between <strong>the</strong>m is<br />

a secondary feature acquired late in development. It has<br />

always been considered that <strong>the</strong> first stage in <strong>the</strong> evolution<br />

<strong>of</strong> <strong>the</strong> Metazoa was a colonial Pro<strong>to</strong>zoon, i. e. a mass <strong>of</strong> perfectly<br />

isolated unicellular org<strong>an</strong>isms, derived by complete division<br />

from a single cell. Now while I do not wish <strong>to</strong> exalt <strong>the</strong> facts<br />

<strong>of</strong> <strong>the</strong> cleavage <strong>an</strong>d early development <strong>of</strong> Peripatus <strong>to</strong> a position<br />

<strong>of</strong> undue import<strong>an</strong>ce, or <strong>to</strong> maintain that <strong>of</strong> <strong>the</strong>mselves<br />

<strong>the</strong>y are sufficient <strong>to</strong> destroy this conception <strong>of</strong> <strong>the</strong> origin <strong>an</strong>d

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