A Revision of the Old World Species (Lepidoptera, Pieridae)

2 Osamu Yata
in space 2 than in space 3; apical patch on forewing underside large, quadrate and
entirely dark chocolate; a spot in discoidal cell; vein-dots on underside not conjointed
with black anticiliary line; uncus moderately long and arched dorsally, with uncal
projection fairly developed.
Description: Male. Upperside'. Ground colour lemon yellow. Forewing
black costal border usually broad with its inner margin sharply defined; black distal
border generally broad, with its inner edge oblique and irregular from costa to vein 4,
more or less angled in the midway, obtuse-angled at vein 4, more deeply excavated in
space 2 than in space 3, almost perpendicular to basal margin or inclined slightly
towards base in spaces la and lb+c; black basal border undeveloped; discocellular
spot absent; fringe black. Hindwing black distal border usually somewhat narrow,
tapering near apex and tornus, with its inner edge usually somewhat diffused and
weakly zigzag-shaped, sometimes projected along each vein; anal border unde
veloped; fringe black, sometimes mixed with yellow. Basal portions of both wings
narrowly blackish. Underside'. Ground colour slightly paler than on upperside.
Forewing apical patch large, quadrate and entirely dark and chocolate with luster; a
3-shaped spot in discoidal cell; discocellular marking represented by an irregular
slender ring, covering more than half of the discocellular vein; tornal spot always
developed; sex-brand usually pale salmon pink, long and narrow, ending slightly
before a point of origin of vein 2; small vein-dots conjointed with a catenate black
anticiliary line; fringe black but mixed with yellow. Hindwing with a series of
submarginal spots in spaces lb+c to 8, arranged in an irregular zigzag-line, and a
submarginal spot in spaces 7 and 8 usually comma-shaped and directed to midway
between submarginal spots and discocellular spot; circular subbasal spot usually
present each in spaces lb+ c and 7 and in the middle of discoidal cell, but a spot in
cell often disappearing; a minute basal spot absent; discocellular spot almost same as
in forewing but larger; small vein-dots appearing in various degree, but not con
jointed with a prominent black marginal streak; fringe yellow, but mixed with black.
Ultraviolet reflectance on upperside: Structurally reflective on yellow area except for
tornal to anal region of hindwing, but diffused distally, appearing bright-white in
UV-photos (PI. 21 (3, upper) in Part II).
Forewing somewhat angulate at apex; distal margin slightly convex. Hindwing
slightly arched in the basal half of costal margin; distal margin evenly rounded; vein
7 usually stalked with vein 6, mdc less than 1/3 length of Idc. Antenna somewhat less
than half the length of forewing, black and white-checkered, except on the posterodorsal
surface and a few apical segments, club cylindrical. Thorax and abdomen
yellow, much darkened above, clothed with black and yellow hairs on thorax and
base of abdomen, a black longitudinal line appearing along the lateral margin of
abdominal terga.
Forewing length: 19.0-23.5 mm.
Female. Similar to male, but differing as follows. Ground colour pale green-