A Revision of the Old World Species (Lepidoptera, Pieridae)

30 Osamu Yata
Fig. 9. Female genitalia of Eurema puella puella (Boisduval, 1832) from Papua New Guinea.
A: Female genitalia (ventral). B: Bursa copulatrix (ventral). C: Signum (anterior).
D: Lateral hollow (lateral). E: Papilla analis (lateral).
ventromedially, obtuse-angled on its anterolateral corner in ventral aspect; longitu
dinal groove weakly sclerotized, broad and deep, but somewhat narrowed medially;
banks of longitudinal groove rather developed, distinctly stippled around ostium
bursae. V-shaped wall undeveloped. Ostium bursae opening at the anterior 1/3 of
genital plate. Ductus bursae about the length of cervix bursae, membranous but
weakly sclerotized on anterior 1/4. Signum moderate in size, with many spines.
Eighth abdominal tergum longitudinally very short; apophysis anterioris nearly
straight, nearly as long as apophysis posterioris, with a prominent protuberance at
the dorsal margin of proximal portion. Papilla analis elongate, bearing a short
apical lobe and swollen bare-region.
Variation: In contrast to E. Candida the present species shows little geographical
variation in wing markings. Seasonal variation is unknown.
Taxonomic remarks: This species had long been considered as conspecific with E.
Candida. However, I raise puella to specific rank here and include subsp. diotima and
papuana from New Guinea, and subsp. octogesa from Waigeo Is. in E. Candida based
mainly on the male genitalia.
Relationship: This species is closely related to E. Candida and forms undoubtedly