A Revision of the Old World Species (Lepidoptera, Pieridae)

28 Osamu Yata
Eurema puella (Boisduval, 1832)
[Pis. 11, 13-15]
Xanthidia puella Boisduval, 1832: 60, pi. 2, fig. 8. (Dutch New Guinea)
Terias papuana Butler, 1898: 60. (Port Moresby)
Diagnosis: Underside of both wings with broad and almost uniform black
distal borders; in hindwing black anal border always absent and black distal borders
usually broad; uncal projection narrow, almost triangular in posterior aspect.
Description: Male. Upperside: Ground colour bright yellow. Forewing
black costal border fairly broad with its inner margin sharply defined; black distal
border generally broad, especially near apex, with its inner edge sharply defined and
almost uniform as a whole, somewhat irregular from costa to vein 4, sometimes very
weakly concaved in spaces 2 and 3, almost perpendicular to basal margin in spaces
la and lb+c; black basal border undeveloped; discocellular spot absent;fringe black.
Hindwing black distal border generally broad, usually tapering near apex and tornus,
with its inner edge almost uniform, but usually very weakly concaved in each space;
anal border undeveloped; fringe black. Basal portions of both wings narrowly
blackish, especially on forewing. Underside: Ground colour slightly paler than on
upperside. Forewing with black distal border almost the same as on upperside;
spotted marking entirely absent; sex-brand pale brown, long and narrow, ending
slightly before a point oforigin ofvein 2; fringe black. Hindwing with black distal
border almost the same as on upperside; spotted makings entirely absent; fringe
black. Ultraviolet reflectance on upperside: Structurally reflective on yellow area
except for tornal toanal region ofhindwing, appearing bright grey in UV-photos (PI.
21 (7, upper) in Part II).
Forewing somewhat angulate at apex; distal margin slightly convex. Hindwing
slightly arched in the basal half ofcostal margin; distal margin evenly rounded; vein
7usually stalked with vein 6, mdc less than 1/3 length oflie. Antenna somewhat less
than half the length of forewing, black, white-checkered on ventral surface, club
cylindrical. Thorax and abdomen usually yellow, much darkened above, clothed
with black and yellow hairs on thorax and base ofabdomen, a black longitudinal line
sometimes appearing along the lateral margin ofabdominal terga.
Forewing length: 18.5-25.0 mm.
Female. Similar to male, but differing as follows. Upperside: Ground colour
usually pure-white. Forewing black costal border narrower, with its inner edge
somewhat diffused; black distal border somewhat broader, with its inner edge more
or less diffused. Hindwing black distal border sometimes broader, with its inner
edge slightly diffused; anal border undeveloped. Underside: Ground colour always
pale yellow except in spaces la and lb+c on forewing. Ultraviolet reflectance on
upperside: Much reflective on ground colour, appearing entirely white in UV-photos
(PI. 21 (7, lower) in Part II).