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Research Highlights of the CIMMYT Wheat Program 1999-2000

Research Highlights of the CIMMYT Wheat Program 1999-2000

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in ECSA rely heavily on <strong>CIMMYT</strong> wheat<br />

germplasm (i.e., 75% <strong>of</strong> released cultivars are<br />

derived directly from <strong>CIMMYT</strong> germplasm).<br />

Regarding germplasm requirements, South Africa<br />

is unique in ECSA, with spring and winter types<br />

grown, as well as F 1<br />

hybrids. Of <strong>the</strong> 71 wheat and<br />

triticale cultivars released in South Africa since<br />

1980, 40% involved <strong>CIMMYT</strong>-derived germplasm,<br />

21% were <strong>of</strong> unknown origin (i.e., <strong>the</strong> parentage <strong>of</strong><br />

commercial F 1<br />

hybrids), with <strong>the</strong> remaining<br />

releases—primarily winter habit types—utilizing<br />

non-<strong>CIMMYT</strong> germplasm.<br />

Diseases Prevalent in ECSA<br />

In recent years, yellow rust has supplanted stem<br />

rust as <strong>the</strong> rust pathogen <strong>of</strong> primary concern in<br />

ECSA. Over <strong>the</strong> past ten years, single step<br />

mutations have resulted in <strong>the</strong> evolution <strong>of</strong> new<br />

virulences in <strong>the</strong> yellow rust pathogen in Kenya,<br />

demonstrating <strong>the</strong> danger <strong>of</strong> breeding for racespecific<br />

resistance. From 1969 to 1989, at least 19<br />

races were identified. In August 1996, yellow rust<br />

was observed for <strong>the</strong> first time in South Africa, and<br />

within a year, <strong>the</strong> disease had spread to most wheat<br />

producing areas in <strong>the</strong> country. Infected grass<br />

species serving as accessory hosts were also<br />

observed in <strong>the</strong> Western Cape and Eastern Free<br />

State, with “6E16” identified as <strong>the</strong> originating<br />

race. MWIRNET/RSA is currently sponsoring a<br />

germplasm shuttle breeding program whereby<br />

South African wheat germplasm is screened for<br />

resistance to yellow rust in a location in<br />

southwestern Uganda considered to be a global<br />

hot-spot for this pathogen.<br />

<strong>CIMMYT</strong> is also supporting distribution <strong>of</strong> a Sub-<br />

Saharan <strong>Wheat</strong> Disease Monitoring Nursery. This<br />

nursery is intended to validate regional varietal<br />

disease performance while monitoring potential<br />

performance criteria important for farmers and<br />

NARS cooperators throughout East, Central, and<br />

Sou<strong>the</strong>rn Africa.<br />

Septoria nodorum, S. tritici and S. avenae f.sp.<br />

triticeae, loose smut, scab, root rot, take-all and<br />

Helminthosporium spp. cause significant yield losses<br />

in ECSA. Cereal aphids (Schizaphis graminum,<br />

Rhopalosiphum padi, R. maidis, Sitobion avenae and<br />

Diuraphis noxia) can also be major pests <strong>of</strong> wheat in<br />

this region, and several species serve as vectors in<br />

<strong>the</strong> transmission <strong>of</strong> barley yellow dwarf virus.<br />

Yield losses <strong>of</strong> up to 47% due to BYDV have been<br />

reported in Kenya. Yield losses <strong>of</strong> >60% due to D.<br />

noxia have been reported in South Africa, and, at<br />

present, all bread wheat cultivars grown in <strong>the</strong><br />

summer rainfall zone <strong>of</strong> RSA require genetic<br />

resistance to this pest. MWIRNET/RSA is currently<br />

supporting research to identify an indicator<br />

capable <strong>of</strong> differentiating Dn resistance genes by<br />

assessing differences in aphid feeding behavior.<br />

Industrial Quality<br />

The free-market, global economy promotes <strong>the</strong><br />

production and consumption <strong>of</strong> <strong>the</strong> lowest cost<br />

products. Throughout ECSA, however, limited and<br />

unscientific information (i.e., market biased) is<br />

available on <strong>the</strong> relative milling and baking quality<br />

<strong>of</strong> nationally produced wheat. ECAMAW and<br />

MWIRNET have promoted Regional Industrial<br />

Quality Cooperative Testing Trials to enable<br />

regional wheat breeders, producers, processors,<br />

and grain marketing organizations to better<br />

understand <strong>the</strong> inherent milling, baking, and<br />

nutritional quality aspects <strong>of</strong> advanced germplasm<br />

and cultivars.<br />

Agronomic <strong>Research</strong><br />

A special feature <strong>of</strong> <strong>the</strong> ECAMAW collaboration is<br />

agronomic research targeted at farm-level problems<br />

that must be resolved to promote long-term<br />

agricultural productivity in <strong>the</strong> region. <strong>Wheat</strong> crop<br />

production recommendations that have recently<br />

been provided to extension services due to<br />

ECAMAW research activities include:<br />

• The semidwarf bread wheat cultivars HAR604<br />

(“Galama”) and HAR1685 (“Kubsa”) recently<br />

released in Ethiopia exhibit <strong>the</strong> highest<br />

productivity and pr<strong>of</strong>itability across a range <strong>of</strong><br />

crop management systems, and result in <strong>the</strong><br />

highest marginal rates <strong>of</strong> return under increased<br />

input levels. The pr<strong>of</strong>itability <strong>of</strong> bread wheat<br />

production in <strong>the</strong> central highlands <strong>of</strong> Ethiopia can<br />

be markedly increased by focusing <strong>the</strong> application<br />

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