<strong>The</strong> <strong>Important</strong> <strong>Bird</strong> <strong>Areas</strong> <strong>of</strong> <strong>Florida</strong>: 2000–2002 – Pranty – 2-Jul-02 22Brown-headed Nuthatch, and Summer Tanager, but others such as Hairy Woodpecker, Red-cockadedWoodpecker, and Bachman's Sparrow do not. For information on Scrubby Flatwoods, see the section onscrub.Sandhills are mixed forests <strong>of</strong> oaks and pines growing on well-drained sandy soils. Many sandhills wereformerly longleaf pine forests that now are dominated by ♦turkey oak (Quercus laevis) and ♦bluejackoak (Q. incana) following clear-cutting <strong>of</strong> the pines and decades <strong>of</strong> subsequent fire exclusion. Somesandhills still retain the open, grassy structure <strong>of</strong> the former flatwoods, while others now are dense forests<strong>of</strong> oaks. Extensive sandhills occur in the western Panhandle and the west-central Peninsula; two sitesknown for their sandhills are Eglin Air Force Base and Withlacoochee State Forest. Many sandhills arebeing restored to flatwoods through removal <strong>of</strong> oaks and an increase in fire frequency. Southern RidgeSandhill is a particular plant community endemic to the Lake Wales Ridge, which runs through theinterior <strong>of</strong> central <strong>Florida</strong>. <strong>The</strong> oaks are composed <strong>of</strong> scrub species and the endemic ♦scrub hickory(Carya floridana) <strong>of</strong>ten is conspicuous. Characteristic breeding birds <strong>of</strong> sandhills depend upon the extent<strong>of</strong> oak/pine and shrub/grass coverages, and may include the Cooper's Hawk, Red-tailed Hawk,“Southeastern” American Kestrel, Northern Bobwhite, Common Ground-Dove, Great Horned Owl,Common Nighthawk, Red-cockaded Woodpecker, Great Crested Flycatcher, Yellow-throated Vireo,Blue Jay, Tufted Titmouse, Brown-headed Nuthatch, Eastern Bluebird, Yellow-throated Warbler, PineWarbler, Common Yellowthroat, Summer Tanager, Eastern Towhee, and Bachman's Sparrow.Hammocks are forests <strong>of</strong> hardwoods (e.g., oaks, hickories, and magnolia) that occur throughout <strong>Florida</strong>.Temperate Hammocks are common along the northern border with Alabama and Georgia, and extendspottily into the central Peninsula, primarily the western half. Hammocks in northern <strong>Florida</strong> containsome <strong>of</strong> the most diverse forests in the eastern United States. Those along the Apalachicola River containplants and animals disjunct from their primary range in the Appalachian Mountains, and these hammockscontain a few endemic trees such as the ♦<strong>Florida</strong> torreya (Torreya taxifolia) and ♦<strong>Florida</strong> yew (Taxusfloridana). ♦Live oaks (Quercus virginiana) and ♦laurel oaks (Q. laurifolia) are abundant in hammocks.Because many oaks in <strong>Florida</strong> are nearly evergreen, hammocks are shaded year-round; as a result, theunderstory <strong>of</strong>ten is extremely sparse and the ground is covered with leaf litter. Fires do not normallyoccur in hammocks. Except in extreme northern <strong>Florida</strong>, ♦cabbage palms (Sabal palmetto)—<strong>Florida</strong>'sstate “tree” (palms aren't trees; they're cycads)—also occur in hammocks, and many hammocks in theprairies north and west <strong>of</strong> Lake Okeechobee are composed entirely <strong>of</strong> cabbage palms. Some examples <strong>of</strong>temperate hammocks occur at Tall Timbers Research Station north <strong>of</strong> Tallahassee, San FelascoHammock Preserve State Park near Gainesville, and Highlands Hammock State Park near Sebring.Maritime Hammocks are temperate hammocks along or close to the Atlantic coast. <strong>The</strong>y <strong>of</strong>ten aresculpted by sea breezes, and are low in stature. Characteristic breeding birds <strong>of</strong> temperate and maritimehammocks include the Cooper's Hawk, Red-shouldered Hawk, Wild Turkey, Yellow-billed Cuckoo,Barred Owl, Red-bellied Woodpecker, Downy Woodpecker, Great Crested Flycatcher, White-eyedVireo, Yellow-throated Vireo, Red-eyed Vireo, Carolina Chickadee, Northern Parula, and SummerTanager. Tropical Hammocks are limited to southern <strong>Florida</strong>, and are composed <strong>of</strong> evergreen trees,shrubs, and palms largely <strong>of</strong> West Indian affinity, such as ♦gumbo-limbo (Bursera simaruba), ♦pigeonplum (Coccoloba diversifolia), ♦false tamarind (Lysiloma latisiliquum), ♦false mastic (Mastichodendronfoetidissimum), and ♦strangler fig (Ficus aurea). Trees and palms are covered with orchids andbromeliads, and ferns carpet the ground. Two subspecies <strong>of</strong> mammals are endemic to tropical hammocks<strong>of</strong> the mainline Keys. Royal Palm Hammock and Mahogany Hammock, both in Everglades <strong>National</strong>Park, as well as the hammocks on Key Largo, are typical examples. <strong>The</strong> diversity <strong>of</strong> avian speciesbreeding in tropical hammocks is quite limited, but several <strong>of</strong> these species occur nowhere else in theUnited States. Characteristic breeding birds include the White-crowned Pigeon, Mangrove Cuckoo, GreatCrested Flycatcher, Gray Kingbird, White-eyed Vireo, and Black-whiskered Vireo. West Indian birds
<strong>The</strong> <strong>Important</strong> <strong>Bird</strong> <strong>Areas</strong> <strong>of</strong> <strong>Florida</strong>: 2000–2002 – Pranty – 2-Jul-02 23that stray to <strong>Florida</strong>, such as the Zenaida Dove, Cuban Pewee, La Sagra’s Flycatcher, Thick-billed Vireo,and Western Spindalis, typically are found in tropical hammocks.Scrub is <strong>Florida</strong>'s oldest plant community, and was formerly common throughout the Peninsula duringperiods <strong>of</strong> lower sea levels and drier climates. Today, scrub is restricted to areas <strong>of</strong> excessively welldrainedsoils, which now occur largely on sand ridges that represent earlier shorelines created duringperiods <strong>of</strong> higher sea levels. Scrub occurs on ridge systems throughout the Peninsula, and along the Gulfcoast in the Panhandle, west barely into Alabama. Xeric Oak Scrub is an early successional form <strong>of</strong>scrub, with numerous patches <strong>of</strong> bare sand, and where vegetation is kept low from intense fires that occurperhaps every eight to 20 years. Because scrub is <strong>Florida</strong>'s oldest plant community and was <strong>of</strong>ten isolatedfrom other habitats by large expanses <strong>of</strong> water, scrub flora and fauna have a high degree <strong>of</strong> endemism; atleast 40% <strong>of</strong> oak scrub species are endemic. <strong>Florida</strong>'s only endemic bird species—the <strong>Florida</strong> Scrub-Jay—is restricted to xeric oak scrub <strong>of</strong> the Peninsula, as are several other vertebrates such as the♦<strong>Florida</strong> scrub lizard (Sceloporus woodi), ♦sand skink (Neoseps reynoldsi), and ♦<strong>Florida</strong> mouse(Podomys floridanus). In the absence <strong>of</strong> fire, xeric oak scrub succeeds to oak hammocks or sand pinescrub, and scrub endemics decline in numbers, potentially to extirpation, unless fire is returned to thecommunity.<strong>The</strong> Lake Wales Ridge is the oldest ridge system in <strong>Florida</strong>, and contains nearly all the oak scrubendemics. <strong>The</strong> endemic ♦scrub oak (Quercus inopina) is common in most interior scrubs, but is notfound along coastal ridges; other oak species are ♦sand live oak (Quercus geminata), ♦myrtle oak(Quercus myrtifolia) and ♦Chapman’s oak (Quercus chapmanii). ♦<strong>Florida</strong> rosemary (Ceratiolaericoides) is a common evergreen shrub <strong>of</strong> many scrublands in <strong>Florida</strong>. Because scrub soils are welldrained,much habitat has been destroyed by the citrus industry, while other scrublands have beeneliminated by commercial and residential development. By the early 1990s, it was estimated that about85% <strong>of</strong> Lake Wales Ridge scrub had been destroyed, and most <strong>of</strong> the remainder would almost certainlysuffer the same fate unless public acquisition activities were initiated quickly. As a result, several Stateand Federal agencies and non-pr<strong>of</strong>it groups partnered to purchase the most significant patches <strong>of</strong> xericoak scrub that remained. Preservation <strong>of</strong> Lake Wales Ridge habitats has been the State's highest priorityfor several years, and as a result, many <strong>of</strong> the most significant scrublands have been preserved. <strong>The</strong> firstscrub preserve—Archbold Biological Station, established in 1941—remains one <strong>of</strong> the most impressiveand most diverse scrub sites in <strong>Florida</strong> (and the world). <strong>The</strong> Merritt Island–Cape Canaveral complex alsocontains large amounts <strong>of</strong> scrub habitats that are undergoing much-needed restoration after a long period<strong>of</strong> fire exclusion. Another State concern is the scrub ridges <strong>of</strong> mainland Brevard County, which are undersevere threat from residential development. Characteristic breeding birds <strong>of</strong> xeric oak scrub are few butinclude the Northern Bobwhite, Mourning Dove, Common Ground-Dove, Common Nighthawk, WhiteeyedVireo, <strong>Florida</strong> Scrub-Jay, Northern Mockingbird, Common Yellowthroat, and Eastern Towhee.Scrubby Flatwoods contain a canopy <strong>of</strong> longleaf or slash pines and a sparse to extensiveunderstory <strong>of</strong> scrub oaks. <strong>The</strong>y occur widely in the central Peninsula. +Abrahamson and Hartnett (1990)note that some <strong>of</strong> scrubby flatwoods may be artifacts <strong>of</strong> previous logging and fire exclusion activities.For a startling and graphic example <strong>of</strong> how quickly and completely oaks can invade pine flatwoods in theabsence <strong>of</strong> fire, see the photographs in +Myers and Ewell (1990: 192) <strong>of</strong> the same landscape at ArchboldBiological Station in 1929 and 1988, after 60 years <strong>of</strong> fire exclusion. Scrubby flatwoods typically containa mixture <strong>of</strong> flatwoods and xeric oak scrub species.<strong>The</strong> third scrub community is Sand Pine Scrub, a forested habitat dominated by ♦sand pines(Pinus clausa). Because most scrubs formerly burned more frequently than at present, sand pine scrubprobably was less common historically than it is at present. Indeed, sand pine forests are the only naturalhabitat that increased in extent during the 20 th century +(Kautz 1993). As noted above, all scrub-endemicspecies decline as the oaks and pines increase in coverage and height, so sand pine scrub supports anentirely different avifauna than does xeric oak scrub. <strong>The</strong> largest patch <strong>of</strong> sand pine scrub in the worldoccurs in and around Ocala <strong>National</strong> Forest. Other patches <strong>of</strong> sand pine scrub occur along the Panhandle
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