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Bryological Monograph An annotated checklist of the mosses of ...

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MOSSES OF EUROPE AND MACARONESIA 233139. Molecular phylogenies <strong>of</strong> Pottiaceae by Werner et al.(2004a, 2005b) place <strong>the</strong> European species in <strong>the</strong> threegroups recognized here: Merceyoideae (Scopelophila),Trichostomoideae (including Splachnobryum,Ephemeraceae, Pleuroweiseae sensu Corley et al.,1981, and some Barbula species, e.g. B. bolleana),and Pottioideae (including Pottieae, Cinclidotus andBarbuleae sensu Corley et al., 1981). Timmiella doesnot belong in Pottiaceae but is listed here until a betterplace can be found for it. Likewise, those Barbulaspecies that belong in Trichostomoideae have beenretained in Barbula until <strong>the</strong>ir correct generic positioncan be ascertained.140. Crumia latifolia (Kindb.) W.B.Sch<strong>of</strong>ield is knownfrom <strong>the</strong> Caucasus but outside Europe.141. <strong>An</strong>oectangium handelii is known from <strong>the</strong> Crimea. It isdescribed by Zander & Weber (2005), correcting anearlier report by Zander (1977) who had treated it as asynonym <strong>of</strong> Molendoa sendtneriana.142. Ephemerum hibernicum was described from Ireland byHolyoak & Bryan (2005).143. Holyoak has examined much material <strong>of</strong> Ephemerumserratum and E. minutissimum, and finds that in bothspecies occasional plants have leaves with a weaknerve. He has not seen <strong>the</strong> type <strong>of</strong> E. serratum var.praecox, which could be a synonym <strong>of</strong> ei<strong>the</strong>r <strong>of</strong> <strong>the</strong>setaxa.144. Ephemerum spinulosum has been found in Ireland and<strong>the</strong> Basque region <strong>of</strong> Spain (Holyoak, 2001; Infante &Heras, 2005).145. Author citation for Eucladium verticillatum follows <strong>the</strong>typification by Ochyra & Zijlstra (2005).146. Gymnostomum mosis (Lorentz) Jur. & Milde wasreported from Spain by Martínez-Sánchez, Ros &Guerra (1991), but later Cano, Ros & Guerra (1994)described <strong>the</strong> new species Gymnostomum lanceolatumbased on this material.147. Gymnostomum aeruginosum var. obscurum wasdescribed by Guerra (2004).148. Spelling <strong>of</strong> species epi<strong>the</strong>t in Hymenostylium recurvirostrumfollows Hedwig (1801) not Crosby et al.(1999).149. Pleurochaete malacophylla (Müll.Hal.) Broth. is listedfor Turkey by Kürschner & Erdağ (2005).150. Splachnobryum delicatulum is treated by Arts (2001) asa synonym <strong>of</strong> S. obtusum. Splachnobryum obtusum wasnot included in <strong>the</strong> main list <strong>of</strong> Corley et al. (1981),because <strong>the</strong>se authors considered that its occurrenceswere not sufficiently natural. However, it occurs in <strong>the</strong>open in Hungary and <strong>the</strong> Azores, and is accordinglylisted here.151. Tortella arctica (Arnold) Crundw. & Nyholm has beenreported from Svalbard, but <strong>the</strong> records refer toTrichostomum arcticum (Frisvoll & Elvebakk,1996).152. Tortella alpicola was reported new for Europe byOtnyukova et al. (2004).153. Tortella bambergeri was not recognized as a goodspecies by Corley et al. (1981), but is generallyaccepted by bryologists in central Europe and alsoby us.154. Tortella densa is treated as a variety T. inclinata var.densa, following Eckel (1998).155. Tortella limbata has been refound on <strong>the</strong> CanaryIslands. Dirkse et al. (1993) compared <strong>the</strong>ir collectionwith <strong>the</strong> holotype and noted <strong>the</strong> distinctive denticulateleaf border, composed <strong>of</strong> long narrow cells, runningnearly to <strong>the</strong> leaf apex.156. Tortella limosella is known only from <strong>the</strong> Scottishtype-locality, and has not been refound since itsdiscovery in 1906.157. Tortella cirrifolia was synonymized with T. nitida bySjögren (2001).158. Molecular studies by Werner et al. (2005b) show that<strong>the</strong> genus Trichostomum as listed here is heterogeneous,with T. brachydontium and T. crispulum nestedwithin Weissia and T. triumphans very close toPottiopsis caespitosa. Trichostomum connivens(Lindb. ex Broth.) Paris is recorded from <strong>the</strong>Caucasus but is not known from Europe.Trichostomum contortum (Kunze) Sérgio is probablya synonym <strong>of</strong> T. brachydontium, although Sérgio(1985) thought that it was a good species.159. For synonymy <strong>of</strong> Hyophila treleasei, see Sollman(1993).160. Weissia leptocarpa Schimp. ex Besch. is treated bySérgio & Carvalho (2003) as a doubtful species, but isaccepted for Turkey by Kürschner & Erdağ (2005), inspite <strong>of</strong> <strong>the</strong> fact that Frey & Kürschner (1991) hadtreated it as doubtful and illegitimate.161. Weissia mittenii (Bruch & Schimp.) Mitt. is omitted,because it is designated above as a hybridW.6mittenii (Bruch & Schimp.) Mitt. emend.A.J.E.Sm. Smith (pers. comm. to Hill) notes that W.mittenii has malformed capsules and abnormal spores.Following Werner et al. (2005b), Astomum is includedin Weissia.162. A molecular study by Werner et al. (2004b) supports<strong>the</strong> species status <strong>of</strong> Weissia wimmeriana.163. Acaulon mediterraneum is treated by many authors asa variety <strong>of</strong> A. muticum, to which it is undoubtedlyclose.164. Aloina humilis was described from <strong>the</strong> Canary Islandsby Gallego et al. (1998).165. Aloina obliquifolia is accorded species rank followingGallego et al. (1999).166. The Macaronesian Barbula semilimbata Dixon &Luisier was listed by Eggers (1982) but is a poorlyknown taxon, not recorded since it was described.167. Barbula convoluta var. sardoa is treated by some atspecies rank. Frahm & Ahmed (2004a) suggested <strong>the</strong>name B. sardoa. However, <strong>the</strong> epi<strong>the</strong>t sardoa is correctat <strong>the</strong> varietal level but superfluous at <strong>the</strong> species level.

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