Bryological Monograph An annotated checklist of the mosses of ...

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MOSSES OF EUROPE AND MACARONESIA 241339. Warnstorfia procera is recognized as a species followingHedenäs (1993b).340. The circumscription of Leskeaceae and Hypnaceaehas been challenged by Gardiner et al. (2005). Werecognize the merit of their proposals, which wouldnecessitate substantial nomenclatural change. Weprefer not to make the changes until a more generaloverview of families in Hypnales is available.341. Lescuraea secunda is reported from European Russiain the Urals (Ignatov & Ignatova, 2004).342. Lindbergia brachyptera, known from the RussianCaucasus, was included by Podpěra (1954) butexcluded by Corley et al. (1981) because theCaucasus was treated by them as part of Asia.343. Pseudoleskea was reinstated as a genus distinct fromLescuraea by Corley & Crundwell (1991), on thebasis of peristome differences emphasized by Crum& Anderson (1981). The peristome characters nowseem less important, and Ignatov & Ignatova (2004)and Söderström (1998) included Pseudoleskea inLescuraea. A conservative position is adoptedhere, pending more detailed analysis by molecularmethods.344. Pseudoleskea brachyclados (Schwägr.) Kindb., originallydescribed from Austria, is listed for Turkey byKürschner & Erdağ (2005). According to Podpěra(1954) it is a variety of P. radicosa (but if so it shouldhave priority at species rank). Other Europeanauthors have treated it as a variety or synonym of P.incurvata.345. We follow Wilson & Norris (1989) in includingLeskeella (Limpr.) Loeske 1903 in PseudoleskeellaKindb. 1897.346. Leskeella incrassata type specimen has been examinedby Ignatov; it is very close to Pseudoleskeella nervosaand, if recognized as a species, will prove to becommon in southern Europe. For the time being thetwo are treated as synonyms.347. Pseudoleskeella rupestris was lectotypified and synonymizedwith P. sibirica by Hedenäs & Söderström(1991).348. In Thuidiaceae, the generic concepts of Touw (2001a)are adopted here. In particular Pelekium Mitt. (1868)emend. Touw is treated as a synonym of Cyrtohypnum(Hampe) Hampe & Lorentz (1869).349. Pelekium atlanticum was described from Madeira asThuidium atlanticum (Hedenäs, 1991). The namePelekium atlanticum (Hedenäs) Touw appears inTouw (2001a), p. 191, but he failed to validate thename by a combination, commenting that P. atlanticumis doubtfully distinct from the American P.muricatulum (Hampe) Touw. On the other hand,Touw was not prepared to treat P. atlanticum as asynonym of P. muricatulum. Hedenäs makes thenecessary combination above.350. For the synonymy of Cyrto-hypnum montei withThuidiopsis sparsa, see Touw (2001a, p. 205). Touwnotes that T. sparsa is known only from a former parkarea in Madeira, and suggests that it is an alien,introduced with ornamental plants.351. Thuidium assimile has priority over T. philibertii, basedon the large-scale revision of Touw (2001b). As notedby Touw, T. assimile belongs to a complex of poorlydefined taxa including T. delicatulum. Observations byMrs C.T. van Dorp on European specimens supportthe recognition of two weakly distinguished species,rather than treating T. assimile as a variety of T.delicatulum.352. Subfamilial and generic concepts in Brachytheciaceaemostly follow Ignatov & Huttunen (2002).These authors overlooked the names HelicodontioideaeM.Fleisch. and Eurhynchioideae Milde, andsuperfluously described these subfamilies asRhynchostegielloideae and Rhynchostegioideae,respectively.353. For the taxonomic status of Thamnium cossyrenseBott. var. cossyrense and var. melitense Bott. seeMastracci (2001).354. Platyhypnidium grolleanum was described from CzechRepublic by Ochyra & Bednarek-Ochyra (1999).355. Platyhypnidium mutatum was described by Ochyra &Vanderpoorten (1999); molecular studies did not finddifferences between this species and P. riparioides(Stech & Frahm, 1999). Frahm (2005a) asserts that itis a somatic mutant of P. riparioides.356. Platyhypnidium torrenticola was described asGradsteinia torrenticola from the Canary Islands byOchyra, Schmidt & Bultmann (1998) and subsequentlytransferred to the genus Platyhypnidium.357. The status of Rhynchostegium arcticum was reevaluatedby Ignatov & Huttunen (2002).358. Rhynchostegium surrectum was synonymized with R.confertum by Hedenäs (1992b).359. Brachythecium cardotii was synonymized withRhynchostegium megapolitanum by Hedenäs (1992b).360. Barbella strongylensis was transferred to Rhynchostegiumby Buck & Privitera (1999).361. Clasmatodon parvulus (Hampe) Sull. is excluded.Heras, Infante & Buck (2006) have found that everySpanish specimen named as Clasmatodon wasPseudoleskeella tectorum. An old German record from1851 is poorly localized (Müller, 2004) and is unlikelyto be C. parvulus.362. Nobregaea latinervis is a morphologically very peculiarspecies known from a single locality in Madeira; it wasdescribed by Hedenäs (1992b).363. Synonymy of Helicodontium capillare follows CortiniPedrotti (2006 [2005]), who indicates that a specimenof the supposedly endemic H. italicum had beenidentified as H. capillare by W.R. Buck.364. Rhynchostegiella bourgaeana is treated by Dirkse &Bouman (1995a) as an endemic of the Canary Islands.On the other hand, Corley et al. (1991) and Hedenäs(1992b) regarded it as a synonym of R. tenella.
242 M. O. HILL ET AL.365. The status of Rhynchostegiella litorea and its differentiationfrom R. tenella are discussed by Dirkse &Bouman (1995a).366. Rhynchostegiella macilenta, omitted by Corley et al.(1981) and Corley & Crundwell (1991), is treated as afull species and illustrated by Dirkse & Bouman(1995a).367. Rhynchostegiella teesdalei and R. jacquinii weresynonymized with R. teneriffae by Dirkse & Bouman(1995a).368. According to Ignatov & Huttunen (2002),Rhynchostegiella tenuicaulis does not belong toBrachytheciaceae and is more likely a member ofAmblystegiaceae, but more studies are needed.369. Rhynchostegiella trichophylla was described from theCanary Islands by Dirkse & Bouman (1995a).370. Molecular data do not confirm the traditional view onthe identity of European Bryhnia scabrida and NorthAmerican B. novae-angliae, so the resurrection of theformer name was suggested by Ignatov & Huttunen(2002).371. Popov et al. (2000) showed that Myuroclada maximowicziiwas reported from Europe on the basis of amisidentification and excluded it from the Europeanflora, but later Virchenko & Babenko (2001) found itas an alien plant in the grounds of the UniversityBotanical Garden, Rostov-na-Donu, southern Russia.372. The status of Brachythecium glaciale var. dovrenseneeds further study. The taxon may deserve recognitionas a species.373. Brachythecium umbilicatum Jur. & Milde is listed forTurkey by Kürschner & Erdağ (2005).374. Brachythecium ryanii was synonymized with B.campestre by Hedenäs (1996).375. Brachythecium rotaeanum and B. capillaceum aretreated as synonyms, as in most recent Europeanchecklists. However, their identity needs confirmation.376. Brachythecium coruscum was recognized as a synonymof B. groenlandicum by Nyholm (1965) but the namewas omitted by Corley et al. (1981).377. Brachythecium erythrorrhizon subsp. asiaticum wasdescribed from the Altai by Ignatov (1998), and isknown from the Urals.378. A typographic error in Wijk et al. (1962) makes itseem that Hypnum thedenii (Schimp.) C.Hartm. (1854)(not Hartm. 1845 as indicated) was published beforeBrachythecium thedenii Schimp. (1853).379. Robinson & Ignatov (1997) showed thatBrachythecium laetum is the correct name for thespecies often called B. oxycladon. In eastern NorthAmerica this species is very common. It is reportedfrom various parts of Europe, but confirmation of theidentity of European plants and the American one isneeded.380. Brachythecium percurrens was described from Madeiraby Hedenäs (1992a). Unpublished molecular data byVanderpoorten et al. and Huttunen et al. suggest thatit belongs to subfamily Helicodontioideae, and may bea member of Oxyrrhynchium.381. Brachythecium rutabulum var. atlanticum wasdescribed from Madeira by Hedenäs (1992a).382. Brachythecium appleyardiae is a synonym ofScleropodium cespitans (Blockeel et al., 2005).383. Brachythecium salteri was synonymized with B. dieckiiby Hedenäs (1993a).384. The combinations Brachytheciastrum fendleri and B.olympicum were made by Vanderpoorten et al. (2005).385. There is still much uncertainty about the correctcircumscription of Hypnaceae and related families.The classification adopted here is provisional, and weare well aware that it will be changed in future.386. The placement of Hypnum lindbergii and H. pratensein Calliergonella and Breidleria, respectively, followsHedenäs (1990).387. The circumscription of Campylophyllum is that ofHedenäs (1997).388. Campylophyllum hispidulum (Brid.) Hedenäs is anAmerican species not found in Europe (Crundwell &Nyholm, 1962).389. Ctenidium molluscum is a very variable species, forwhich numerous infraspecific taxa have beendescribed. The following varieties are recognized inmore than one European country: var. condensatum(Schimp.) E.Britton, var. molluscum, and var. robustumBoulay. The genetic basis of this variationrequires further elucidation.390. Hypnum cupressiforme has such a complex patternof variation that the delimitation of infraspecifictaxa remains highly controversial. The varieties inthe main list are those recognized in current checklistsof at least four European countries. Other names,some of which may have priority over more popularones, are var. brevisetum Schimp., var. imbricatumBoulay, var. julaceum Brid., var. tectorum Brid.,var. tenue Hook. & Taylor, and var. tectorum (Brid.)J.-P.Frahm.391. Hypnum heseleri was described from Saarland inGermany and two localities in the Netherlands(Ando & Higuchi, 1994). van Zanten & Hofman(1994) grew spores from capsules of H. heseleri anddemonstrated that the progeny were about 50% H.heseleri and 50% H. cupressiforme. Allozyme-electrophoresisprovided further convincing evidence that H.heseleri is a locally derived mutant of H. cupressiforme.Accordingly, H. heseleri is treated as a varietyof H. cupressiforme in a new combination, madeabove.392. Hypnum aemulans Breidl. ex Limpr., originallydescribed from Austria, is treated as a goodspecies by Ando (1994) but is listed withoutcomment as a synonym of H. hamulosum byEuropean authors (Koperski et al., 2000; Ulvinen,Syrjänen & Anttila, 2002). European occurrences needfurther investigation.
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