Bryological Monograph An annotated checklist of the mosses of ...

MOSSES OF EUROPE AND MACARONESIA 239autoicous specimens and the differences he reportedfrom B. lonchocaulon (other than in sexuality) appearinconstant in material from Britain.295. Bryum bimum may merit species rank since it differsfrom B. pseudotriquetrum in being synoicous ratherthan autoicous, perhaps in having twice as manychromosomes (Smith, 2004), and in average dimensionsof the leaf lamina cells and thickness of their cellwalls (Demaret & Empain, 1985; Zolotov, 2000).However, they apparently do not differ much, if at all,in ranges, frequency or ecology, and preliminarymolecular data (Hedenäs, pers. comm. to Holyoak)suggest they are very close.296. Bryum salinum differs from B. archangelicum in only afew characters; a few intermediate specimens areknown.297. Bryum schleicheri author citation follows Geissler(1985).298. Wijk et al. (1959) cited the original description ofBryum torquescens as Bruch ex De Not., Syllab.Musc., no. 163, 1838. Syed (1973) found this referenceto be wrong and he discounted ‘Fl. Sard. Exs., 1828’,given by other authors because he suspected it is anomen nudum. The original description is thereforetaken as that by Bruch & Schimper (Bryol. Eur., 4 p.119, pl. 20, 1839 (fasc. 6–9, Mon., p. 49, pl. 20)) partlybecause this was the earliest description found by Syedand partly because they refer to it as ‘Diese neue Art’.299. Arts, Crundwell & Whitehouse (1995) showed thatBryum pyriferum described from Tenerife is conspecificwith B. valparaisense, which was originallydescribed from Chile but is now also known from N.America, Africa and Portugal.300. Hedwig (1801) used the spelling zierii twice, but thename Bryum zierii commemorates John Zier as waspointed out by Crum & Anderson (1981). More recentauthors who amend it to zieri are therefore correctingit according to Article 60.11 of the International Code(Greuter et al., 2000).301. Touw (1984) gave reasons for rejecting the nameRhodobryum spathulatum in favour of R. ontariense.302. The large family Mniaceae of Goffinet & Buck (2004)is here divided following Koponen (1988) into thethree families traditionally included in Mniaceae andthe remainder, for which the only available name isMielichhoferiaceae.303. Mielichhoferia elongata is often treated as a synonymor variety of M. mielichhoferiana, but according toShaw (2000a), it is in reality a cryptic species.304. Pohlia ramannii Warnst. is recorded from the territorythat was formerly in Finland and now in Russia(Söderström, 1998). It is not mentioned by Ignatov &Afonina (1992) and is treated here as doubtful.Webera luisieri Dixon and W. maderensis Dixon &Luisier, listed by Eggers (1982), are obscure taxawhose identity is uncertain.305. Sections of Pohlia are based on Shaw (1984), exceptthat we follow Smith (2004) in placing P. crudoides insection Pohlia and P. flexuosa in section Cacodon.306. Pohlia bolanderi was found in the Sierra Nevada, newfor Europe (Rams et al., 2004).307. Pohlia ambigua was described from Austria butremains a poorly known taxon. According to Shaw(1982) it is a form of P. elongata. It is autoicous and istherefore assigned to var. acuminata.308. Species limits in the Pohlia nutans group requirefurther elucidation. Köckinger, Kučera & Stebel(2005) studied P. schimperi and established that theonly really stable character separating it from P.nutans is its red coloration; they accordingly gave itsubspecies rank and synonymized it with P. nutansvar. purpurascens. Pohlia marchica Osterwald wasdescribed from near Berlin and subsequently collectedon Bornholm. It has not been found for many years,and is signified by Nyholm (1993) as very close to P.schimperi. Pohlia turonensis J.M.Couderc & Guédès,nom. inval. was described from France. It is a closerelative of P. nutans and has not subsequently beenvalidated.309. Pohlia saprophila, a mainly Asian species, is reportedfrom the Komi Republic, Russia (Zheleznova, 1994).It may prove to be a dioicous race of P. elongata, butneeds further study.310. For the synonymy of Pohlia flexuosa see Townsend(1995).311. Smith (2004) recognized two varieties of Pohliaflexuosa in Europe, namely var. flexuosa and var.pseudomuyldermansii nom. nud. (P. muyldermansiivar. pseudomuyldermansii Arts, Nordhorn-Richter &A.J.E.Sm.). Townsend (1995) had noted that there iscontinuous variation in bulbil morphology in Nepaland India, and argued that the varieties shouldtherefore be treated as synonyms. He hypothesizedthat there might be only two clones in Europe, withvar. muyldermansii possibly a recent introduction.However Smith (pers. comm. to Hill) is of the opinionthat in Europe there has been differentiation into twomorphologically and ecologically distinct groups ofplants. In view of the distribution of var. pseudomuyldermansii(Austria, British Isles and Switzerland)it seems unlikely that the populations concernedrepresent a single clone. The habitat of the varietiesappears to be markedly different in Europe. For thesereasons, Smith (above) makes the new combination P.flexuosa var. pseudomuyldermansii.312. Pohlia tundrae is reported from Central Europe (Düll,1991, 1992; Müller, 2004).313. Mnium lycopodioides is treated here as a full synonymof M. ambiguum following Koponen (1994).314. There are a number of records of Trachycystisussuriensis in the Russian Caucasus, e.g. Kharzivovet al. (2004).