Bryological Monograph An annotated checklist of the mosses of ...

MOSSES OF EUROPE AND MACARONESIA 243393. Hypnum holmenii was described from Canada,Greenland and Finland (Ando, 1994), and has beenfound in the Arkhangelsk Oblast of Russia (Ignatov &Ignatova, 2004).394. Hypnum subimponens is reported from EuropeanRussia by Afonina (2004).395. Vesicularia sphaerocarpa (A.Jaeger) Broth. wasreported from Malta by Reimers in 1934 on the basisof a 19th century collection by Schweinfurth. Thespecimen (B) was destroyed in the Second World War,but Reimers in Bizot & Potier de la Varde (1952) wasconfident that it belonged to the taxon described bythese authors as V. reimersiana. According to Corleyet al. (1981), V. sphaerocarpa is a synonym of V.galerulata (Duby) Broth. Düll (1985) listed theMaltese plant as V. reimersiana, but also asserted thatV. galerulata had been found in mainland Italy.Corley & Crundwell (1991) accordingly added V.reimersiana to the list, retaining V. galerulata.However, no species of Vesicularia is listed for Italyby Cortini Pedrotti (2006 [2005]). It seems likely thatV. galerulata has never been found in Europe.396. Smith (2004) treated Heterocladium flaccidum as aspecies, but provided only a nomen nudum. He hasmade the necessary new combination above for thischecklist, noting (pers. comm. to Hill) that consultationwith a number of British field bryologists revealedthat, with one exception, none had encounteredintermediates between it and H. heteropterum.397. Heterocladium wulfsbergii is discussed in detail byCrundwell & Smith (2000).398. Iwatsukiella leucotricha was listed for Europe byPodpěra (1954) under the name Habrodon leucotrichus(Mitt.) Perss., with a locality in the southern Urals. Itwas omitted without comment by Corley et al.(1981), but is indicated from the European side ofthe Urals by Dierssen (2001) and Ignatov & Ignatova(2004).399. Alpine and arctic forms of Hylocomium splendens havebeen variously treated. Var. alpinum Schlieph. exLimpr. is mapped by Söderström (1996), but isdoubted and treated as possibly a synonym of var.obtusifolium (Geh.) Paris by Koperski et al. (2000).Var. obtusifolium is commonly treated as a synonymof H. alaskanum (Lesq. & James) Austin, whosedistinguishing characters have been shown by Ross etal. (2001) to be mainly under environmental ratherthan genetic control. It is not clear whether any ofthese forms is sufficiently distinct genetically todeserve taxonomic recognition.400. Herzogiella adscendens (Lindb.) Z.Iwats. & W.B.Schofieldhas been reported from Svalbard (Corley &Crundwell, 1991; Düll, 1985), but the recordwas subsequently rejected (Frisvoll & Elvebakk,1996).401. Myurella julacea var. ciliata (Chal.) Ochyra &Bednarek-Ochyra (syn. M. julacea var. scabrifoliaLindb. ex Limpr.) is treated as distinct in someEuropean countries, but intergrades completely withvar. julacea. It is not recognized here.402. Orthothecium chryseon is incorrectly listed as O.chryseum by Crosby et al. (1999). It was described asHypnum chryseon by Schwägrichen (1804).403. Plagiothecium curvifolium is sometimes treated as avariety (in Europe) or synonym (in North America)(Ireland, 2003) of P. laetum, to which it is undoubtedlyvery close. As a variety, its correct name is P. laetumvar. secundum.404. Plagiothecium denticulatum var. undulatum is treatedin many European lists as a species, P. ruthei, but inNorth America merely as a synonym of P. denticulatum(Ireland, 2003). It is not clearly distinct from var.denticulatum, and we follow Ignatov & Ignatova(2004), Koperski et al. (2000) and Smith (2004) intreating it as a variety.405. Plagiothecium noricum was described from thePinzgau, Austria and was thought by Molendo(but not Limpricht) to intergrade with P. neckeroideum.Grims (1999) listed it but noted that Düll(1992) had thought it a weak species, possiblyonly a variety of P. neckeroideum. It is not recognizedhere.406. Plagiothecium succulentum is a difficult taxon,intergrading on the one hand with P. nemorale(Hemeric, 1989) and on the other with P. cavifolium(Nyholm, 1965). It is also very close to P. platyphyllum.It is retained here, because it cannot satisfactorilybe subordinated to any of these species withouttreating the whole group as a single, rather-broadspecies.407. Plagiothecium svalbardense was described by Frisvollin Frisvoll & Elvebakk (1996).408. Pseudotaxiphyllum laetevirens was raised to speciesrank by Hedenäs (1992a).409. Entodon challengeri occurs in European Russia(Ignatov & Ignatova, 2004) and is the correct namefor the plant formerly known as E. compressus(Iwatsuki & Tan, 2001).410. According to Ochyra & Ireland (2004), Isopterygiumtenerum is absent from Europe but present in sub-Saharan Africa. However, Schumacker in Stewart(1995), Cortini Pedrotti (2006 [2005]) and Gabrielet al. (2005) list it for Italy and the Azores.Isopterygium bottinii was excluded from Sematophyllumby Guerra & Gallego (2005), and is includedhere in I. tenerum.411. Hageniella micans, previously known from Europeand North America, was synonymized with H.pacifica Broth., which occurs in China and Taiwan(Tan & Jia, 1999).412. Sematophyllum adnatum was found in Italy, where it isthought to be an introduction from North America(Brusa in Blockeel et al., 2000).