Bryological Monograph An annotated checklist of the mosses of ...

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MOSSES OF EUROPE AND MACARONESIA 235pseudohandelii as a synonym of the non-European S.caninervis var. pseudodesertorum (Vondr.)M.T.Gallego, which is found in western Asia, includingTurkey.206. For the status of Syntrichia caninervis var. abranchesiirefer to Gallego et al. (2002a).207. Syntrichia caninervis var. astrakhanica was describedby Ignatov, Ignatova & Suragina (2002).208. Syntrichia glabra was described by Frahm & Gallego(2001).209. Following Gallego, Cano & Guerra (2004), noinfraspecific taxa are recognized within Syntrichialaevipila, which is synonymized with S. pagorum,Tortula laevipilaeformis and T. saccardoana.210. Syntrichia minor was recognized at species rank byGallego et al. (2000).211. According to Ochyra (1994), Tortula intermedia DeNot. 1838 is a synonym of Tortella humilis, makingTortula intermedia (Brid.) Berk. 1863 an illegitimatehomonym.212. According to Gallego (2005), Tortula papillosa var.meridionalis Warnst. should not be recognized as adistinct variety.213. According to Gallego (2005), Syntrichia princeps var.parnassica (Schiffn.) Podp. should not be recognizedas a distinct variety.214. Author citation of Syntrichia ruralis var. ruraliformisfollows Gallego (2005).215. Syntrichia subpapillosissima was recognized at speciesrank by Gallego et al. (2002b). These authors treatedTortula ruralis var. submamillosa W.A.Kramer as asynonym.216. Tortula buyssonii (H.Philib.) Broth., from theAuvergne, France, is a little-known taxon that hasapparently not been found since the 19th century.217. Tortula caucasica Lindb. (Pottia caucasica (Lindb.)Paris) is known from Georgia but not from Europe.Tortula grandiretis Broth. is listed for Turkey byKürschner & Erdağ (2005).218. Tortula ampliretis, described from the Canary Islandsin a paper by Crundwell, During & Long (1978), wasinexplicably not mentioned by Corley et al. (1981).219. Tortula bolanderi and T. inermis should be in Tortulanot Syntrichia (Werner et al., 2003a).220. Tortula israelis, described from Spain as T. muralisvar. baetica, was raised to species rank by Guerra &Ros in Guerra, Ros & Carrión (1992) and subsequentlysynonymized with T. israelis by Cano, Guerra& Ros (1996).221. According to Cano (2004a), European plants attributedto Crossidium seriatum Crum & Steere should bereferred to Tortula brevissima.222. The synonymy and nomenclature of Tortula hoppeanafollow Ochyra (2004a).223. For the synonymy of Desmatodon meridionalis seeSérgio & Granzow de la Cerda (2001).224. According to Sérgio & Carvalho (2003), the typematerial of Pottia cuneifolia corresponds to P. pallida.Therefore, P. pallida and Tortula zoddae, which is anew name for P. cuneifolia in Tortula, are included inthe synonymy of T. pallida.225. Zander (1993) provided the new name Tortularhodonia for Desmatodon wilczekii, because the epithetwilczekii is not available in Tortula.226. Cano, Werner & Guerra (2005) raised Tortula subulatavar. angustata with bistratose border to species rank, onthe basis of T. angustata Lindb., and provided the newname T. schimperi, because the epithet angustata is notavailable at species rank in Tortula. The other varietiesof T. subulata were sunk in a single variable species.227. According to Cano (2004a, b), Tortula marginatasubsp. limbata is a synonym of T. solmsii.228. Tetraplodon urceolatus is a good species which hasbeen misunderstood in Scandinavia, where it has beenconfused with forms of T. mnioides.229. Orthotrichum urnaceum Müll.Hal. was signified forEurope by Lewinsky-Haapasaari (1995) but thelocality is in Armenia.230. Synonymy of Orthotrichum limprichtii followsLewinsky-Haapasaari in Nyholm (1998).231. Orthotrichum casasianum is reported only from northernSpain (Mazimpaka et al., 1999).232. Orthotrichum crenulatum was reported as O.flowersii Vitt from the French Alps (Boudier &Pierrot, 1992).233. Orthotrichum handiense is an endemic fromFuerteventura, Canary Islands (Lara et al., 1999a;Lara, Garilleti & Mazimpaka, 2003).234. Orthotrichum hispanicum was described from Spain byLara, Garilleti & Mazimpaka (2000).235. Orthotrichum macrocephalum was described fromSpain by Lara, Garilleti & Mazimpaka (1994).236. For synonymy of Orthotrichum lewinskyae see Draperet al. (2003).237. Orthotrichum schimperi is recognized as a species inaccordance with Cortini & Lara in Cortini Pedrotti(2001).238. For synonymy of Orthotrichum rogeri var. defluens,see Garilleti, Lara & Mazimpaka (2002).239. For synonymy of Orthotrichum australe, see Lewinsky(1993).240. Orthotrichum vittii was described from Spain by Laraet al. (1999b).241. For synonymy of Orthotrichum arcangelianum seeLara et al. (2002) and Lewinsky (1993).242. Orthotrichum shawii has been reinstated as a distinctspecies (Mazimpaka et al., 2000).243. Orthotrichum ibericum was described from Spain andPortugal by Lara & Mazimpaka (1993).244. For the synonymy of Orthotrichum caucasicum, seeLewinsky (1993).245. We have excluded Orthotrichum elegans Schwägr. as aEuropean species because Nordic material named as
236 M. O. HILL ET AL.O. speciosum var. elegans (Schwägr.) Warnst. isreferable to O. speciosum. For differences betweenthese two taxa see Vitt & Darigo (1997).246. Orthotrichum speciosum var. brevisetum is known fromSpain & Sicily; see Draper et al. (2003).247. Orthotrichum tortidontium was described fromMorocco and Spain by Lara, Garilleti & Mazimpaka(1996).248. Orthotrichum vladikavkanum was described from theRussian Caucasus by Venturi (1887). It was listed inPodpěra’s (1954) Conspectus, but the Caucasus wasassigned to Asia by Corley et al. (1981). For thepresent enumeration, the Russian part of the Causasusis now again included in Europe.249. Corley et al. (1981) included Ulota macrospora in U.rehmannii, but the two species are distinguishedhere, following Boudier & Pierrot (1996) and Sauer(1998).250. The inclusion of Zygodon forsteri in Codonoblepharonis controversial (Goffinet and Vitt, 1998; Matcham &O’Shea, 2005). In our opinion it is best placed inZygodon. Casas et al. (1985) considered Z. madeirensisto be close to Z. forsteri; it is treated here as asynonym following Düll (1992). Further revision isneeded to assess its identity.251. Zygodon sibiricus was reported from the Europeanside of the Urals by Ignatov & Ignatova (2003).252. Zygodon stirtonii is recognized at species rank followingSmith (2004).253. Hedwigia ciliata var. viridis Bruch & Schimp. is listedby Casas (1991), but Hedenäs (1994) was unable totrace the type specimen, so it is uncertain whether thevariety belongs to H. ciliata or H. stellata.254. Hedwigia ciliata var. leucophaea was illustrated andmapped by Hedenäs (1994).255. Hedwigia stellata was described from Sweden byHedenäs (1994).256. Anacolia laevisphaera is reported from Europe in anabstract by Quandt & Frahm (2004). It is a tropicalmontane species and German specimens from theMosel region had been incorrectly identified asBartramia stricta.257. Anacolia menziesii was found new to Europe in theprovinces of Almería and Granada in southern Spain(García-Zamora et al., 1998). Its distinctness from A.webbii was confirmed by molecular data (Werner etal., 2003b).258. Bartramia pomiformis var. elongata Turner and B.pomiformis var. heteromalla (Brid.) Müll.Hal. aretreated with great scepticism by Nyholm (1998), andvar. elongata is treated as a synonym of var.pomiformis by Smith (2004). These varieties areaccordingly excluded from the checklist.259. Nomenclature for Bartramia section Pyridium (sectionVaginella Müll.Hal., nom. illeg.) follows Ochyra et al.(2003).260. Bartramia breviseta was originally described fromFinnmark, but was treated as a synonym of B.ithyphylla in many European checklists. Fransén(2004) has demonstrated that it is distinct, differing,for example, in the lack of an inner peristome.261. Bartramia ithyphylla var. strigosa (Wahlenb.) Hartm.was mapped by Söderström (1996). However, Fransén(2004) treats it as a synonym of var. ithyphylla, and itis accordingly not recognized here.262. Philonotis marchica var. laxa, described fromSwitzerland and Germany as P. laxa Limpr., is listedby Casas (1991) but is no longer recognized in Germanand Swiss lists.263. Philonotis uncinata (Schwägr.) Brid. is listed for theAzores by Gabriel et al. (2005).264. Molecular data now leave no doubt that the generaAnomobryum, Brachymenium, Plagiobryum andRhodobryum cannot be maintained with the circumscriptionused by Corley et al. (1981) since most areingroups within Bryum sensu lato (e.g. Pedersen, Cox& Hedenäs, 2003; Pedersen & Hedenäs, 2005). Themolecular data also show that there have beenmultiple instances of convergent evolution of morphologicalcharacters in Bryaceae, so that classificationsbased exclusively on the morphologicalcharacters currently known are unsound. Hence, forexample, the genus Rosulabryum proposed by Spence(1996) is not adopted here because molecular dataimply it is polyphyletic (Pedersen et al., 2003). A starthas been made on subdividing Bryum into segregategenera (e.g. Imbribryum N.Pedersen) on the basisof molecular data from multiple loci used in combinationwith morphological data (Pedersen, 2005;Pedersen & Hedenäs, 2005), but their treatmentcannot yet be applied to European species as awhole because molecular data are not availablefor many species. The generic treatment of Corleyet al. (1981) is therefore used here in an interimarrangement.265. Loeske (1932 [1933]) long ago interpreted Anomobryumleptostomoides as fertile A. concinnatum, andnumerous fertile specimens examined by Holyoakfrom its type-locality at Mont Dore (France)consistently show the gametophytic characters ofconcinnatum. This is therefore the only Europeanlocality at which A. concinnatum has been foundwith capsules, and it is significant because sporophytesof A. leptostomoides differ from those of A.julaceum in the erect not inclined to cernuouscapsules and their greatly reduced peristome, asdescribed in detail by Shaw & Fife (1984).Furthermore, similar capsules have been describedfrom Michigan (USA), again from plants withgametophytes indistinguishable from those of A.concinnatum. Shaw & Fife (1984) were ‘inclined torecognize A. leptostomoides at the specific level’ buthesitated to do so because they thought that the
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