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Sclerophyton<br />
Arthoniales Roccellaceae<br />
15<br />
Sclerophyton is a genus confined to the tropical and oceanic areas <strong>of</strong> the world.<br />
But it is possible that this genus is found in all parts <strong>of</strong> the world below an altitude<br />
<strong>of</strong> 1500m, with a mean temperature over 5° C and an annual precipitation over<br />
500mm. In Sclerophyton, the highest species diversity occurs in South and Central<br />
America and tropical Asia. Temperate Europe, Central America, Australia and<br />
Southeast Asia have their own species. Out <strong>of</strong> all known species <strong>of</strong> Sclerophyton,<br />
only four are considered as widespread.<br />
Typically, corticolous species <strong>of</strong> this genus are found on palm beaches, coastal<br />
scrublands and in woodlands. Saxicolous species grow on shaded, <strong>of</strong>ten<br />
overhanging rocks. Some species grow in tropical rainforests. Most species are<br />
distributed within a small area. All species <strong>of</strong> Sclerophyton grow on nutrient poor<br />
to nutrient rich, acid to neutral or inert substrates, such as smooth, acidic bark,<br />
hard wood and sclerophyll leaves (hard leaves with short internodes).<br />
Ramalinaceae<br />
Fr<br />
VC<br />
Ramalina usnea<br />
Description The fruticose thallus is corticolous, pendulous, long, yellowish<br />
brown, and branched with ribbon-like branches that are spirally twisted, and are<br />
<strong>of</strong>ten cracked. Pseudocyphellae are linear. Soralia are absent. Medulla is solid.<br />
Apothecia are not found in <strong>Sri</strong> <strong>Lanka</strong>n specimens.<br />
Status Very common species.<br />
Habitat Mostly restricted to humid high elevation forest above 1400 m.<br />
Locations Horton Plains and Nuwara Eliya.<br />
Though this genus is mostly found in dry habitats, the species included here is<br />
observed in an upper montane area above 1300 m, along the Pattipola road in<br />
the Nuwara Eliya District. This is the first species <strong>of</strong> this genus to be recorded<br />
in <strong>Sri</strong> <strong>Lanka</strong> (Weerakoon & Aptroot unpublished). It is possible that many other<br />
species <strong>of</strong> this genus could be present in <strong>Sri</strong> <strong>Lanka</strong>, as other genera <strong>of</strong> the family<br />
Roccellaceae are frequently recorded.<br />
Apothecia round to elongate or asteroid, immersed, generally in groups or lines,<br />
disc exposed, pink or pale to dark brown to black, <strong>of</strong>ten pruinose, margins white<br />
to grey, <strong>of</strong>ten fusing to form stromata (a thick compact wad <strong>of</strong> hyphae, <strong>of</strong>ten<br />
covering or supporting a group <strong>of</strong> ascomata); Asci: 8-spored; Spores: colourless,<br />
clavate, septate (3-10) or muriform.<br />
Chemistry Thallus UV- or UV+ yellow , PD+ yellow or PD+ orange , K+ Yellow or<br />
K+ yellow turning red, C+ red.<br />
Roccellaceae<br />
Oceanic areas: Pertaining to a climate characterized by mild, wet winters, cool,<br />
moist summers, and frequent fogs.<br />
146 <strong>Fascinating</strong> <strong>Lichens</strong> <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong><br />
<strong>Fascinating</strong> <strong>Lichens</strong> <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong> 147