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Hymenogaster and related genera.pdf - MykoWeb

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630 ANNALS OF THE MISSOURI BOTANICAL GARDEN<br />

[VOL. 21<br />

conical; basidia -1-4-spored; spores colored, usually some shade<br />

of brown, ochre, or almost black at maturity, ovoid, obovoid,<br />

ellipsoid, citriform, fusiform, or lanceolate, with or without<br />

apiculus, with or without persistent pedicel, surface smooth<br />

to verrucose, rugose, alveolate, or reticulate, with or without<br />

a more or less wrinkled utricle.<br />

The peridium <strong>and</strong> tramal tissues of the gleba are little dif-<br />

ferentiated in- the young stages. In some species they never<br />

undergo further differentiation, while in others the outer<br />

hyphae become larger <strong>and</strong> form a coarse stupose outer layer<br />

which gradually merges into the gleba. The cavities develop<br />

schizogenetically in the upper portion of the fructification,<br />

leaving a large more or less hemispherical to conical sterile base<br />

which persists for a long time, occasionally to maturity. As the<br />

cavities exp<strong>and</strong> above, the lower ones next the sterile base are<br />

stretched longitudinally so that they appear to radiate from<br />

the sterile base. The septa are very broad at first <strong>and</strong> grad-<br />

ually shrink until they are very thin, often fragile, occasionally<br />

scissile. The subhymenium is often pseudoparenchymatous,<br />

the basidia are usually cylindrical <strong>and</strong> so evanescent that they<br />

are rarely observable in mature fructifications. On drying, the<br />

tissues throughout the fructification collapse, <strong>and</strong> thus meas-<br />

urements based on dry material are many times less than those<br />

based on fresh specimens or on those preserved in alcohol.<br />

When a fructification has been preserved in alcohol for a time<br />

<strong>and</strong> then allowed to dry out, the tissues are still more 'collapsed<br />

<strong>and</strong> it is almost impossible to distinguish structures. Since<br />

most of the material available in this work has been dried,<br />

unless otherwise noted the measurements <strong>and</strong> the structures<br />

observable in the dry material have been given in our descrip-<br />

tions <strong>and</strong> keys. One who intends to prepare herbarium ma-<br />

terial of this genus should make careful field notes of color,<br />

odor, <strong>and</strong> size, <strong>and</strong> preserve most of the material in alcohol or<br />

alcohol to which 10 per cent glycerol has been added, drying<br />

one fructification in order to compare with dry material to be<br />

found in other herbaria. Exceptionally fine material received<br />

from Prof. James McMurphy, Stanford University, was pre-

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