42 bearing small gland; smaller glandular dots also rarely present. Filament dark. Ovary, 1.0-1.25 whorls, overlapping posterior stomach chamber. Albumen gland without a pallial component. Capsule gland as long as albumen gland. Genital aperture a subterminal slit without vestibule. Coiled oviduct a short kink followed by small circular coil and broad, horizontal loop. Oviduct and bursal duct join well behind pallial wall. Bursa copulatrix pyriform, posterior end angled, almost as long and broad as albumen gland, with about 65% <strong>of</strong> length posterior to albumen gland. Bursal duct narrow, shallowly embedded in albumen gland proximally, short. Seminal receptacle pouch-like, short, overlapping anterior bursa copulatrix, extending to posterior edge <strong>of</strong> albumen gland. TYPE LOCALITY.—[Snake River at] Homedale, Owyhee County, Idaho. Baker (1964) incorrectly stated that a holotype was selected by Pilsbry. Pilsbry figured two specimens, but did not indicate which if any <strong>of</strong> <strong>the</strong>se is <strong>the</strong> type. Herein I designate a lectotype, ANSP 152677, consisting <strong>of</strong> <strong>the</strong> first <strong>of</strong> <strong>the</strong> two specimens illustrated by Pilsbry. The o<strong>the</strong>r specimen is a paralectotype, ANSP 396960. DISTRIBUTION.—Snake River (main stem), southwestern Idaho. REMARKS.—Relatively few anatomical specimens were available for this animal. Additional details can be gleaned from <strong>the</strong> unpublished report <strong>of</strong> Taylor (1982). MATERIAL EXAMINED.—USNM 874698, gravel bar just upstream from mouth <strong>of</strong> Clover Creek, Elmore County, Idaho (T5S,RllE,SWl/4sec8). <strong>Pyrgulopsis</strong> intermedia (Tryon, 1865) Pomatiopsis intermedia Tryon, 1865:220, pi. 22: fig. 8.—Baker, 1964:173. Bythinella intermedia.—Tryon, 1870:49, pi. 16: fig. 7. Fontelicella (s.s.) intermedia.—Gregg and Taylor, 1965:108.—Taylor, 1985:309-310. Fontelicella intermedia.—Taylor, 1975:104.—Burch, 1982:26, fig. 238.— Turgeonetal., 1988:61. <strong>Pyrgulopsis</strong> intermedia.—Hershler and Thompson, 1987:29. Paludestrina longinqua.—Pilsbry, 1899:122 [in part].—Steams, 1901:284, fig. 2 [in part].—Hannibal, 1912b:186 [in part].—Walker, 1918:138 [in part]. DIAGNOSIS.—Shell ovate-conic, large, narrowly umbilicate. Penial filament and lobe medium length. Penial ornament a penial gland, transverse terminal gland, and ventral gland. DESCRIPTION.—Shell (Figure \la,b) ovate-conic; height, 4.0-5.5 mm; whorls, 4.5-5.0. Earliest protoconch weakly punctate, o<strong>the</strong>rwise near smooth except for a few spiral lines. Teleoconch whorls highly convex; sculpture <strong>of</strong> variably pronounced growth lines. Aperture ovate, adnate or slightly separated from body whorl. Inner lip complete, slightly thickened (sometimes greatly in columellar area); columellar lip slightly reflected. Outer lip slightly prosocline. Umbilicus narrowly rimate, sometimes excavated. Periostracum tanbrown. Operculum (Figure \lc,d) ovate, amber with nuclear region SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY especially dark; nucleus slightly eccentric; dorsal surface frilled. Attachment scar margin broadly thickened all around, region between nucleus and inner edged raised; callus well developed. Central radular tooth (Figure 36c) with moderately indented dorsal edge; face near square; lateral cusps, 4-5; central cusp rounded, considerably broader, slightly longer than laterals; basal cusps, 1, short, with strong dorsal support. Basal process medium width; basal sockets deep. Lateral margins thickened; neck weak-absent. Cephalic tentacles pale except for small black internal pigment patch distal to eyespots. Snout moderate gray. Foot moderate gray, darker along anterior, posterior edges. Opercular lobe dark along sides and anterior edge. Neck lighter than snout, foot. Pallial ro<strong>of</strong>, visceral coil dark, <strong>of</strong>ten near-uniform black. Ctenidial filaments, 25, medium height and width. Osphradium centered slightly posterior to middle <strong>of</strong> ctenidial axis. Kidney with medium-large bulge into pallial cavity; opening white. Stomach caecum pronounced, triangular. Testis, 1.75 whorls, overlapping stomach to edge <strong>of</strong> style sac. Prostate gland with large (25%) pallial section; pallial vas deferens proximally kinked. Penis (Figure 47ft) medium-sized, extending beyond mantle edge; base near-square, with folds along inside edge; filament medium length, tapering; lobe about as long as filament, broad, with little taper. Penial gland covering about two-thirds <strong>of</strong> filament. Terminal gland large, usually transverse, borne along distal edge <strong>of</strong> lobe. Ventral gland stalked, positioned near mid-length. Filament with dark internal pigment. Ovary, 1 whorl, overlapping anterior stomach chamber. Albumen gland without a pallial section. Capsule gland as long as albumen gland. Genital aperture a terminal slit with short vestibule. Coiled oviduct a short posterior oblique twist followed by near horizontal loop, nearly abutting pallial wall anteriorly. Oviduct and bursal duct join anterior to oviduct coil just behind pallial wall. Bursa copulatrix ovoid-pyriform, medium length and width, with about half <strong>of</strong> length posterior to albumen gland. Bursal duct slender, shallowly embedded in albumen gland, medium length. Seminal receptacle pouch-like, short, with short duct, positioned lateral to proximal bursal duct near ventral edge <strong>of</strong> albumen gland. TYPE LOCALITY.—Owyhee River, sou<strong>the</strong>ast Oregon. Type material is labeled Crooked Creek, Owyhee, sou<strong>the</strong>ast Oregon. ANSP 2795 8 [a] contained 4 specimens segregated (by Baker) as types, 1 <strong>of</strong> which was fur<strong>the</strong>r segregated from <strong>the</strong> o<strong>the</strong>r 3. The latter was broken, and <strong>of</strong> gerontic appearance, and did not resemble Tryon's figure although <strong>the</strong> shell measurements were similar to those he gave. I chose ano<strong>the</strong>r, unbroken, adult shell as lectotype, ANSP 27958; paralectotypes, ANSP 396959. DISTRIBUTION.—Sou<strong>the</strong>rn Oregon: Crooked Creek, Owyhee River drainage (Snake basin); Barren Valley (internal drainage), lower Klamath Lake basin; nor<strong>the</strong>ast California: Pit River below Fall River (Taylor, 1985:309).
NUMBER 554 FIGURE 17.—Western <strong>Pyrgulopsis</strong>: a-d, P. intermedia (a, shell, ANSP 27958,3.4 mm; b, shell, USNM 874195, 4.0 mm; c4, opercula, bar = 0.67 mm); e-g, P. isolata (
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- Page 23 and 24: NUMBER 554 17 Pyrgulopsis amargosae
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- Page 27 and 28: NUMBER 554 21 Pratt (1977) recorded
- Page 29 and 30: NUMBER 554 23 DISTRIBUTION.—This
- Page 31 and 32: NUMBER 554 25 nucleus slightly ecce
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- Page 35 and 36: NUMBER 554 29 FIGURE 12.—Western
- Page 37 and 38: NUMBER 554 31 FIGURE 13.—Western
- Page 39 and 40: NUMBER 554 33 ornament an elongate,
- Page 41 and 42: NUMBER 554 35 dium centered slightl
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- Page 51 and 52: NUMBER 554 45 FIGURE 18.—Western
- Page 53 and 54: NUMBER 554 copulatrix. Seminal rece
- Page 55 and 56: NUMBER 554 Pyrgulopsis merriami (Pi
- Page 57 and 58: NUMBER 554 FIGURE 20.—Western Pyr
- Page 59 and 60: NUMBER 554 or lightly pigmented on
- Page 61 and 62: NUMBER 554 weakly frilled. Attachme
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- Page 67 and 68: NUMBER 554 61 FIGURE 24.—Western
- Page 69 and 70: NUMBER 554 63 units; Dg3 small (som
- Page 71 and 72: NUMBER 554 65 Pyrgulopsis simplex H
- Page 73 and 74: NUMBER 554 67 FIGURE 26.—Western
- Page 75 and 76: NUMBER 554 69 DIAGNOSIS.—Shell ov
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- Page 79 and 80: NUMBER 554 FIGURE 28.—Eastern Pyr
- Page 81 and 82: NUMBER 554 75 medium length, obliqu
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- Page 93 and 94: NUMBER 554 87 Pyrgulopsis bryantwal
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NUMBER 554 93 FIGURE 36.—Central
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NUMBER 554 FIGURE 38.—Central rad
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NUMBER 554 97 FIGURE 40.—Central
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NUMBER 554 99 FIGURE 42.—Central
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NUMBER 554 FIGURE 44.—Penes of we
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NUMBER 554 FIGURE 46.—Penes of we
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NUMBER 554 FIGURE 48.—Penes of we
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NUMBER 554 FIGURE 50.—Penes of we
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NUMBER 554 FIGURE 52.—Penes of ea
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Abbott, R.T. 1948. A Potential Snai
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NUMBER 554 Society of Washington, 1
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NUMBER 554 115 gyrus, Gabbia, Hydro