24 SSIITHSONIAN CONTRIBUTIONS TO BOTANY FIGURE 12.4pecies <strong>of</strong> <strong>Parmelina</strong>: a, P. damaziana (Dainazio 1375 in W); b, P. degelii (Degelius A-I82 in Degelius herbarium); c, P. de?iegans (<strong>Hale</strong> 43855); d, P. dissecta (<strong>Hale</strong> 14999); e, P. endoleuca (Weber in <strong>Lichen</strong>es Exsiccati 244 in US): f, P. enorinis (Jellicoe in US). (Scale in mm.)
NUMBER 33 <strong>the</strong> chemical constitutents, P. damaziana is a rare and not well comprehended species. The lectotype was collected on tree branches whereas Lynge’s species was apparently saxicolous. Burchell 1105-06 is tentatively placed here since <strong>the</strong> chemistry, while not clear, is closest to <strong>the</strong> “horrescens” type. Parme- lina damaziana has ra<strong>the</strong>r large spores, just as <strong>the</strong> o<strong>the</strong>r members <strong>of</strong> <strong>the</strong> P. horrescens group. It could be regarded <strong>the</strong>oretically as <strong>the</strong> parent morph for P. howescens, P. schindleria, and P. subfatiscens, but all <strong>of</strong> <strong>the</strong>se vegetative morphs have smaller, more fragile thalli. SPECIXIENS ExAalrNED.-BraLil: Burchell 1105-06 (BM, US). 8. <strong>Parmelina</strong> degelii, new species FIGURE 12h DEscRIPrroN.-Thallus adnatus, corticola, fragilis, pallide viridi-albicans, 6-8 cm latus, lobis plus minusve subirreqularibus, 2-3 mm latis, margine aetate lobulato-dissectis, ciliatis, ciliis usque ad 0.8 mm longis, superne planus vel aetate rugulosus, nitidus, sparse vel modice isidiatus, isidiis primum cylindricis, mox procumbentibus, dorsiventralibus, expansis, ad 0.6 mm longis, cortex superior 12 pm crassus, epicorticatus, stratum gonidiale 12-14 pm crassum, medulla pallide sulfurea, 60-90 pm crassa, cortex inferior paraplectenchymatus, brunneus, 12 p,m crassus, subtus niger, dense rhizinosus, rhizinis nigris, simplicibus, 0.5-09 mm longis. Apo<strong>the</strong>cia ignota. CHEMIsTRY.-Cortex K + yellow (atranorin); medulla more intensively yellow with color tests (zeorin, leucotylic acid, apparently secalonic acid A, and traces <strong>of</strong> o<strong>the</strong>r pigments and terpenes). HoLoTYPE.-Rain forest, Cairns Cove, Queensland, Australia, elevation 600 m, G. Degelius A-I82 (Degelius herbarium; isotype in US). DI~TRIRUTI~N.-A~S traha. REMARKS.-This is <strong>the</strong> only species <strong>of</strong> section Myelochroa that has been discovered in Australia. It is remarkable that such a conspicuous lichen should be collected so recently in Queensland. The terpene chemistry is close to P. aurulenta except for <strong>the</strong> absence <strong>of</strong> one spot (Figure 7). Regardless <strong>of</strong> chemistry, however, it is distinct from all o<strong>the</strong>r species in <strong>the</strong> P. subaurulenta complex because <strong>of</strong> <strong>the</strong> production <strong>of</strong> large lobulate isidia. <strong>Parmelina</strong> degelii is named in honor <strong>of</strong> Dr. Gunnar Degelius, who has enriched our knowledge <strong>of</strong> lichen distribu- tions through his careful collecting efforts in <strong>the</strong> tropics. 9. <strong>Parmelina</strong> denegans FILLRF 12C Parmelzna denegans (Nylander) <strong>Hale</strong>, 1974,482. Parnelta denegans Njlander, 1900 6 [tjpe collection: Ram- podde, Ceylon, Almquzst (H, Njlander herbarium number 35129, lectotype; isolectotype in S)]. DE~CRIPTI~X .-Thallus adnate to closely adnate on bark, light greenish mineral gray, 5-9 cm broad; lobes irregularly branched, sublinear, imbricate, 1-3 mm wide, <strong>the</strong> marginal cilia irregularly dispersed, 0.4-0.7 mm long; upper surface shiny, weakly maculate, plane but soon developing small pustulate areas, <strong>the</strong> pustules eventually breaking open apically and forming coarse soredia; medulla pale orange-salmon, in part K+ purple; lower surface black, densely rhizinate, <strong>the</strong> rhizines black, simple to sparsely branched. Apo<strong>the</strong>cia sessile, 2-3 mm in diameter; spores 8, 5-7 X 7-9 p.m. CHEhmTRY.-Cortex K + yellow (atranorin); medulla more intensively yellow-orange with color tests (zeorin, leucotylin and associated terpenes, secalonic acid A, and possibly anthraquinones). DIsTRIsuTIoN.-India, Sri Lanka, and Sabah. REMARKS.-Except for <strong>the</strong> darker reddish orange pigment in <strong>the</strong> lower part <strong>of</strong> <strong>the</strong> medulla, this species is <strong>the</strong> pustulate morph <strong>of</strong> P. subaurulenta. They have similar lobe configuration and habitats, occurring at about 2000 m elevation in open forests. The lectotype specimen is fertile and has few pustules. The specimens that I collected are heavily pustulate-sorediate by contrast. SPECIMENS ExAM1xFD.-India: Tamil Nadu, Foreau 4128 (Awa3thi herbarium, US), <strong>Hale</strong> 40239, 43636, 43762, 43855. Sri Lanka: Thruaites (UPS). Sabah: <strong>Hale</strong> 29020. 10. <strong>Parmelina</strong> dissecta FIGCJRE 12d. 13 <strong>Parmelina</strong> dissecta (Nylander) <strong>Hale</strong>, 1974:482. Parmelia dissecta h’ylander, 18823451 [type collection: Fon- tainebleau, France, A’ylander (H, Nylander herbarium number 35131, lectotype; isolectotype in PC)]. Parinelia laeuigata var. gracilis f. furfuracea Miiller Argovien- 25
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