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A Monograph of the Lichen Genus Parmelina Hale - Smithsonian ...

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26 SMITHSONIAN CONTRIBUTIONS TO BOTANY<br />

sis, 1888c:529 [type collection: Faxina, Brazil, Puiggari 47<br />

(G, lectotype)].<br />

Parmelia minarum Vainio, 1890:48.<br />

Parmelia amazonica var. hwnotii Hue, 1899:158.<br />

Parmelia puiggari Gyelnik, 1931:288 [based on Parmelia<br />

lamigata var. gracilis f. furfuracea Muller Argoviensis].<br />

Parmelia camtschadalis var. epiphylla Cengia Sambo,<br />

1938:379.<br />

Parmelia hubrichtii Berry, 1941:102.<br />

[For full citation <strong>of</strong> synonymy, see <strong>Hale</strong>, 1971a:6.]<br />

DEScRIPTIoN.-Tha1Ius adnate on bark or rock,<br />

yellowish glaucous to pale greenish mineral gray,<br />

3-7 cm broad: lobes sublinear-elongate, contiguous,<br />

1-3 mm wide, <strong>the</strong> marginal cilia irregularly dis-<br />

persed, mostly simple, to 0.7 mm long: upper sur-<br />

face shiny, emaculate, plane to convex, moderately<br />

to densely isidiate, <strong>the</strong> isidia cylindrical, erect,<br />

<strong>of</strong>ten branched, less than 0.5 mm high; medulla<br />

white; lower surface black, moderately rhizinate,<br />

<strong>the</strong> rhizines black, shiny, simple or sparsely<br />

branched. Apo<strong>the</strong>cia adnate, <strong>the</strong> rim crenate, <strong>the</strong><br />

amphi<strong>the</strong>cium isidiate: 1-4 mm in diameter: spores<br />

8,8-10 X 12-17 pm.<br />

CHEMIsmY.-Cortex K+ yellow (atranorin);<br />

medulla K-, C+ rose, KC+ red, P- gyrophoric<br />

acid with or without associated unknown sub-<br />

stances).<br />

DIsTRIBuTI0N.-Pantemperate and montane pan-<br />

tropical (Figure 13).<br />

REMAws.-This is one <strong>of</strong> <strong>the</strong> most widespread<br />

and commonly collected species in <strong>the</strong> genus,<br />

especially in temperate zones, occurring on a wide<br />

variety <strong>of</strong> substrates in open, secondary forests. In<br />

tropical regions it is strictly montane, usually being<br />

most abundant in cloud forests up to 2300 m eleva-<br />

tion. Morphological variation is wide, even though<br />

<strong>the</strong> basic characters, isidia, ciliate lobes, and gyro-<br />

phoric acid, are constant. The lobes, for example,<br />

are narrow, sublinear, and separate in <strong>the</strong> lectotype<br />

specimen. At <strong>the</strong> o<strong>the</strong>r extreme are plants such as<br />

those collected in Dominica (<strong>Hale</strong>, 1971a) with<br />

broader, contiguous to subimbricate lobes.<br />

FIGURE 13.-Distribution <strong>of</strong> <strong>Parmelina</strong> dissecta based on all available herbarium specimens.

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