Cassiinae pt 1 NY-Botanical_gardens_Vol. 35_1 - Copy.pdf - Antbase

362 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [VQL. 35
genuine C. laevigata, but since his sy^nymy has become generahy accepted the
genuine C. floribunda has dropped out of sight.
The fact remains that there is a senna distinct from C. laevigata that grows
wUd in Mexico and answers exceUently to the plant grown at Madrid in the first
decade of the XIX century and independently described by Gomez Ortega as
Cassia corymbosa (not of Lamarck) and by Cavanihes as C. floribunda. It is,
however, found only in the cornmon range of C. laevigata (=our Senna septem­
trionalis) and S. multiglandulosa, at least sometimes' i^n close association with
both. Moreover it is morphologicaUy intermediate in outline and pubescence of
the leaflets, in the ovule-number, and in the arrangement ofthe seeds, which are
strictly 2-seriate in S. septemtrionalis, stricly 1-seriate in S. multiglandulosa, but
vary from 1-seriate to inegularly displaced in what we here call S. x floribunda.
LocaUties where these three sennas are known to grow close together are on the
peripheral foothills of VaUe de Mexico (numerous coUections), around Zitacuaro
and Temascaltepec in Mexico State (Hinton 6216, 8278, 13546), and along the
San Luis Potosi-Rio Verde highway near Santa Catarina (septemtrionalis:
Roe & Roe 2160, Schubert & Souza 1919; multiglandulosa: Lasseigne
4887; X floribunda: Lasseigne 4886, 4889, HiU & Funk 1784). We feel convinced
that Cassia floribunda is a fertile, self-perpetuating hybrid that probably was
synthesized several times as opportunity arose for exchange of poUen between
the parent species, features of which it incorporates in varying degree. The typus
of Adipera bicolor, for example, has the relatively small elliptic, basaUy asymmetric
leaflets of S. multiglandulosa, but they are glabrous or almost so. In the
typus of A. arsenei the leaflets are larger and densely pubescent, in fact resembling
those of the original C. floribunda which also came from Puebla. Different
strains of the hybrid differ in flower-size, the handsome plant known in gardens
as Cassia herbertiana representing the large-flowered extreme.
117. Senna punoensis Lasseigne, sp. nov., habitu, inflorescentia et legumine S.
multiglandulosae praesimihs ac manifeste affinis, sed foliohs lanceolato-ovatis
(nee oblongo-eUipticis) 7-11 (nee 6-8)-jugis superne glabris
inferne puberuhs (nee lanatis) necnon staminum 2 abaxialium longorum
antheris apice horizontaliter (nee oblique) truncatis diversa. PERU.
Puno, prov. Sandia: along stream among rocks in open, 3200-3450 m,
near Limbani, 14-16.V.1942, R. D. Metcalfe in Goodspeed 30491.—
Holotypus, UC; isotypus, MO.
Potentially arborescent to 6 m but probably flowering when smaher and only
shrubby, the annotinous branches castaneous, the hornotinous, densely leafy
ones with h-stalks and dorsal face of lfts either strigulose with straight subappressed
or pUosulous with loosely spreading-ascending hairs up to 0.2-0.45(-0.75)
mm, the vesture commonly lutescent, the ample crowded fohage strongly bicol­
ored, the hts dull ohvaceous or dark green and glabrous above, palhd beneath,
the axillary racemes equaUing or shorter than the subtending If, forming an immersed
or scarcely emergent corymbiform panicle.
Stipules erect submembranous, palhd or brownish, hnear-lanceolate 2.5-
7.5 X 0.4-1.2 mm, early caducous, absent from mature spms.
Lvs (7-)8-15.5 cm; petiole including discolored but little swollen pulvinus
7-15(-18) mm, not or Uttie longer than the first interfoholar segment of rachis, at
middle 1-1.4 mm diam, the shaUow open ventral sulcus narrowly thick-margined;
rachis 5.5-10.5 cm, hs longer segments 8-14 mm; glands between most pairs or
ah except the distal pair of hts, sessUe or substipitate 0.8-1.3 mm, the ovoid