Cassiinae pt 1 NY-Botanical_gardens_Vol. 35_1 - Copy.pdf - Antbase

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382 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [VOL. 35 13. Blade of staminodes trapeziform (broadest distally); beak of 2 long abaxial anthers 0.7-1.2 mm; local in BeUze and immediately adjoining Guatemala and Mexico. 126j. var. hemirostrata (p. 389). 13. Blade of staminodes inversely deUate; beak of 2 long anthers (l-)1.2-2 mm, at orifice 0.5-0.9 mm; widespread in range indicated at choice 11 above. 126k. var. advena (p. 389). 12. Beak of 2 long abaxial anthers drum-shaped, broader than long; style 6.5-10 mm and long abaxial filaments 15-20 mm; Colombia (if brought higher by plants of s.-w. Venezuelan lowlands with style and filaments barely within Umits of variation indicated at choice 2 above, cf. 1261, var. meticola. 126i. var. pendula (p. 388). 11. Plants of Brazil, Peru and s. through BoUvia to Paraguay and n. Argentina. 14. Adult lfts sharply and closely reticulate beneath or on both faces; BraziUan Planalto s.-ward from near 13°S in Bahia and Goias to n.-e. Argentina, in Brazil extending locaUy to the coast in Parana and Sta. Catarina. 15. Pod subcyhndric, the seeds 2-seriate; common and widespread between 13° and 26°S, but not coastal. 126a. var. glabrata (p. 382). 15. Pod turgid but laterally compressed, the seeds 1-seriate; either coastal in s.-e. BrazU or of n.-e. Argentina. 16. Staminodes narrowly oblanceolate, ± twice as long as wide; ovules ±48-52; coastal s.-e. BrazU (Parana and Sta. Catarina). 126b. var. recondita (p. 384). 16. Staminodes rhomboidal, scarcely longer than wide; ovules 58-98; Misiones and adjoining Corrientes, Argentina. 126c, var. missionum (p. 384). 14. Lfts not sharply reticulate on either face, the tertiary venulation faint or erratic or immersed. 17. Anther of 2 long abaxial stamens 5.5-8 mm, obviously shorter than its filament; widespread. 18. Distribution in the e. foothiUs and interior vaUeys of the Andes from centr. Peru to n.-w. Argentina; petiolar glands between aU but the distal pair of lfts; blade of staminodes either shorter than 2 mm or narrower than 1.6 mm (or both). 19. Longest sepal 8-11 mm and longest petal 14-18(-20) mm; filament of 2 long abaxial stamens 12-18 mm; blade of staminodes oblong 3-4.5 mm; Peru, BoUvia. 126e. var. praeandina (p. 385). 19. Longest sepal 5.5-6 mm and longest petal 10.5-13 mm; filament of 2 long stamens 8-10 mm; blade of staminodes rhombic-suborbicular 1.2-1.5 mm; n.-w. Argentina. 126f. var. eriocarpa (p. 386). 18. Distribution along rivers of n.-e. Brazil in Para, Maranhao and e.ward; petiolar glands at proximal and rarely the second pair of lfts; blade of staminodes amply trapeziform 2.7^ x 2-2.5 mm. 126g. var. indistincta (p. 387). 17. Anther of 2 long abaxial stairiens 13-14 mm, as long or a trifle longer than its filament; local in n.-e. Minas Gerais and Paraiba, Brazil. 126h. var. dolichandra (p. 387). 1. Lfts ovate or ovate-elliptic, broadest weU below middle; local in Amazonian Peru (San Martin). 126s. var. scandens (p. 395). 126a. Senna pendula (WUldenow) var. glabrata (Vogel) Irwin & Barneby, comb. nov. Cassia indecora var. [8] glabrata Vogel, Syn. Gen. Cass. 19. 1837, in statu iUegit. (subordinated to a species containing an earUer proposal).—"In BrasiUa: SeUow legit pr. Ypanema [Sao Paulo]."— Holotypus, tB; neotypus, presumed isotypus, K! Cassia dormiens VeUozo, Fl. Flum. 159. 1825 & Ic. 4: t. 67. 1835.—Lectotypus, the cited p which shows the large fl and biseriate seeds of the var! Cassia coluteoides Colladon, Hist. Casses 102, t. 12. 1816.—". . . culta in hortis . . . v.v. sine fl. in h[orto] m[eo] et s.c. in herbario d. Bouchet."—Lectotypus, an old cult, spm dating back to DeCandoUe's years at MontpelUer, the stock acquired from Lisbon, MPU! Cassia crassisepala Bentham, Linnaea 22: 527. 1849.—". . . [in BrasiUae provincia Minarum generaUum imprimis prope Caldas coUegerat clar. A. RegneU.] 11.76"—Holotypus, Regnell
1982] CASSIINAE—SENNA 383 76 of collection II, K (hb. Benth.)!—Reduced by Bentham, 1870, p. 106 and 1871, p. 525, to C. bicapsularis, sens. lat. Cassia bicapsularis (typical) sensu Bentham, 1870, p. 107, majori ex parte; Burkart in Parodi, Encicl. Argentina Agric. y Jard. 1: 465, fig. 464C (optima!). 1959. Diffuse or assurgent shrubs, in cenado commonly 1-3 m, at margin of gallery forest sarmentose to 5 m, commonly glabrous throughout but the hornotinous stems and lf-stalks sometimes pUosulous and the hts then, or independently, pUosulous dorsally in distal basal angle of midrib, rarely dorsally pilosulous overaU; petiolar gland between proximal, occasionally also the second (-fourth) pair of lfts, these usually 4-6, in many populations 4-5, in some 6-7 pairs, the distal pair mostly 2.5-5.5(-6.5) x 0.9-2.3 cm, their secondary camptodrome veins 7- 12 pairs, the intervenium intricately reticulate; longest sepal (9-)10-14.5(-l5.5) mm; longest petal (16-) 18-26 mm; blade of staminodes typicaUy rhombic-orbic ular to subquadrate and 1.5-2.7(-3) x (l-)1.4-2.2 mm, rarely (n.-ward, transient to var. indistincta) oblong-pandurate and 3.6-4 x 1.6-3 mm; 2 long filaments 13- 18.5 mm, their anther (7-)7.5-l0 x 1.5-2.2 mm, its beak 0.6-1 mm; style 5-8 mm; ovules mostly 70-96, locaUy up to 132; pod subcylindric ±9-16 x (0.9-)l- 1.6 cm; seeds 2-seriate.—Collections: 103.—Fig. 10 (androecium), 12 (pod, seed). Cerrado, disturbed cerradao, margin of gallery forest and about granite or sand stone outcrops, mostly 450-1100 m but ascending to campo rupestre at 1320 and locaUy in Sa. do Espinhago (Sa. da Piedade) to 2000 m, becoming weedy along roadsides and in disturbed woodland, common and widespread over the Brazilian Planalto from centr. Mato Grosso to s.-centr. Bahia, s. to interior Parana and s.centr. Paraguay, reappearing locally on restinga and at margin of wet forest in coastal s. Bahia; cultivated in warm temperate United States and locally naturaUzed in s. peninsular Florida and the Bahamas (Great Abaco).—Fl. in S. America (XI-)I-VI(-VII), the pods maturing slowly and long persisting.—Fedegoso, cassia (Brazil) smd pito-muvero (Paraguay), but these used genericahy for various sennas; caquera (reported by Vellozo for this or a closely alhed form) is not confirmed by any modern collector. This is the handsome large-flowered, longistylous senna that in southeastern Brazil has passed, along with assorted kindred described hereafter, as typical Cassia bicapsularis, a mistake traceable back to Bentham (1870, p. 106-107); and that latterly formed a substantial part of C. pendula var. pendula sensu Lasseigne (ined.). The original Cassia pendula is here regarded as a smaU entity endemic to the eastern cordillera in Colombia at points distant over 2500 km to the northwest of var. glabrata, from which it differs in the simpler, non-reticulate venu lation ofthe leaflets and in the broadly cunehorm or inversely deltate staminodes. Except that it may possibly be cultivated on the Atlantic slope in Rio or Guanabara, var. glabrata is not found along the coast of southeastern BrazU where it is replaced by the closely related, respectively longi- and brevistyled vars. re condita and ambigua, and equaUy different from h in the compressed pod and uniseriate seeds. A pod entirely similar in compression is the principal diagnostic character also of longistylous var. missionum, which replaces var. pendula vi- cariantly along the lower Paraguay river in Misiones, Argentina. Beyond the intravarietal variation accounted for in our description it should be mentioned that the populations found in wet coastal forest chmax and adjoining restinga along the coast of Bahia tend to have relatively ample and thin-textured but nonetheless finely reticulate leaflets. The eariiest collection of this form (Luschnath in Martius 715) contributed to the protologue of C. bicapsularis var. tenuifolia, which lectotypification permhs us to preserve for a quite different Amazonian variety ofthe species.
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•'•mmmmmmfii^'''- '->mmmmmmmm^'
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ADVISORY BOARD MEMOIRS OF THE N E W
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This material is based upon researc
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IV FOREWORD In looking back over th
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DECEMBER 1982 MEMOIRS OF THE NEW YO
Page 13 and 14:
1982] CASSIINAE—INTRODUCTION 1. F
Page 15 and 16:
1982] CASSIINAE—CASSIA Fig. 1. An
Page 17 and 18:
1982] CASSIINAE—CASSIA commonly m
Page 19 and 20:
1982] CASSIINAE—CASSIA C. fistula
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1982] CASSIINAE—CASSIA 11 sulcate
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1982] CASSIINAE—CASSIA 13 i. ser.
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1982] CASSIINAE—CASSIA 15 1.3-2.8
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1982] CASSIINAE—CASSIA 17 modific
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1982] CASSIINAE—CASSIA 19 and is
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1982] CASSIINAE—CASSIA 21 Adult l
Page 33 and 34:
1982] CASSIINAE—CASSIA 23 Fig. 3.
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1982] CASSIINAE—CASSIA 25 cm, the
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1982] CASSIINAE—CASSIA 31 exactly
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1982] CASSIINAE—CASSIA 33 bowl-sh
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1982] CASSIINAE—CASSIA 35 s.-e. V
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Page 49 and 50:
1982] CASSIINAE—CASSIA 39 like) a
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1982] CASSIINAE—CASSIA 41 sometim
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1982] CASSIINAE—CASSIA 43 sylvis:
Page 55 and 56:
1982] CASSIINAE—CASSIA 45 Lvs 13-
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1982] CASSIINAE—CASSIA 49 f produ
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1982] CASSIINAE—CASSIA 51 garden
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1982] CASSIINAE—CASSIA 53 Klug, G
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1982] CASSIINAE—CASSIA 55 Cassia
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1982] CASSIINAE—CASSIA 57 Not see
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1982] CASSIINAE—CASSIA 59 Cassia
Page 71 and 72:
1982] CASSIINAE—CASSIA 61 Opposit
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1982] CASSIINAE—CASSIA 63 of leaf
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1982] CASSIINAE—SENNA 65 neglecta
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1982] CASSIINAE—SENNA 67 Pubescen
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1982] CASSIINAE—SENNA 69 tionary
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1982] CASSIINAE—SENNA 71 zygomorp
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1982] CASSIINAE—SENNA 73 apical s
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1982] CASSIINAE—SENNA 75 alternat
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1982] CASSIINAE—SENNA 77 our clas
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1982] CASSIINAE—SENNA 79 pressed
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1982] CASSIINAE—SENNA 81 technica
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1982] CASSIINAE—SENNA 83 parallel
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1982] CASSIINAE—SENNA 85 18. Pod
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1982] CASSIINAE—SENNA 87 3. Senna
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1982] CASSIINAE—SENNA 89 tertiary
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1982] CASSIINAE—SENNA 91 never so
Page 103 and 104:
1982] CASSIINAE—SENNA 93 3d. Senn
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1982] CASSIINAE—SENNA 95 broad ma
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1982] CASSIINAE—SENNA 97 with ran
Page 109 and 110:
1982] CASSIINAE—SENNA 99 ed. 2, 1
Page 111 and 112:
1982] CASSIINAE—SENNA 101 duction
Page 113 and 114:
1982] CASSIINAE—SENNA 103 broadsi
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1982] CASSIINAE—SENNA 105 4. Blad
Page 117 and 118:
1982] CASSIINAE—SENNA 107 many-fl
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1982] CASSIINAE—SENNA 109 neighbo
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1982] CASSIINAE—SENNA 111 through
Page 123 and 124:
1982] CASSIINAE—SENNA 113 a multi
Page 125 and 126:
1982] CASSIINAE—SENNA 115 Vesture
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1982] CASSIINAE—SENNA 117 those o
Page 129 and 130:
1982] CASSIINAE—SENNA 119 obhquel
Page 131 and 132:
1982] CASSIINAE—SENNA 121 Chamaef
Page 133 and 134:
1982] CASSIINAE—SENNA 123 3 abaxi
Page 135 and 136:
1982] CASSIINAE—SENNA 125 little
Page 137 and 138:
1982] CASSIINAE—SENNA 127 biseria
Page 139 and 140:
1982] CASSIINAE—SENNA 129 Trees o
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1982] CASSIINAE—SENNA 131 Lf-rach
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1982] CASSIINAE—SENNA 133 Strongl
Page 145 and 146:
1982] CASSIINAE—SENNA 135 interca
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1982] CASSIINAE—SENNA 137 cium it
Page 149 and 150:
1982] CASSIINAE—SENNA 139 atively
Page 151 and 152:
1982] CASSIINAE—SENNA 141 pus, da
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1982] CASSIINAE—SENNA 143 affinit
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1982] CASSIINAE—SENNA 145 quangul
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1982] CASSIINAE—SENNA 147 . . ."
Page 159 and 160:
1982] CASSIINAE—SENNA 149 pair of
Page 161 and 162:
1982] CASSIINAE—SENNA 151 Amply l
Page 163 and 164:
1982] CASSIINAE—SENNA 153 of the
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1982] CASSIINAE—SENNA 155 Hithert
Page 167 and 168:
1982] CASSIINAE—SENNA 157 27. Sen
Page 169 and 170:
1982] CASSIINAE—SENNA 159 persal
Page 171 and 172:
1982] CASSIINAE—SENNA 161 Weak di
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1982] CASSIINAE—SENNA 163 ovary d
Page 175 and 176:
1982] CASSIINAE—SENNA 165 Lvs 4-5
Page 177 and 178:
1982] CASSIINAE—SENNA 167 Pod pen
Page 179 and 180:
1982] CASSIINAE—SENNA 169 Fig. 16
Page 181 and 182:
1982] CASSIINAE—SENNA 171 coriace
Page 183 and 184:
1982] CASSIINAE—SENNA 173 petiole
Page 185 and 186:
1982] CASSIINAE—SENNA 175 ±3.5-8
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1982] CASSIINAE—SENNA 177 a membe
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1982] CASSIINAE—SENNA 181 hkewise
Page 193 and 194:
1982] CASSIINAE—SENNA 183 leaflet
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1982] CASSIINAE—SENNA 185 present
Page 197 and 198:
1982] CASSIINAE—SENNA 187 42f. Se
Page 199 and 200:
1982] CASSIINAE—SENNA 189 Lvs bel
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1982] CASSIINAE—SENNA 191 or acut
Page 203 and 204:
1982] CASSIINAE—SENNA 193 Stipule
Page 205 and 206:
1982] CASSIINAE—SENNA 195 leafles
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1^^^] CASSIINAE—SENNA 197 In and
Page 209 and 210:
1982] CASSIINAE—SENNA 199 parenri
Page 211 and 212:
1982] CASSIINAE—SENNA 201 r^jCico
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1982] CASSIINAE—SENNA 207 Chamaef
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1982] CASSIINAE—SENNA 209 dackU i
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1982] CASSIINAE—SENNA 215 Araguai
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1982] CASSIINAE—SENNA 217 Racemes
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1982] CASSIINAE—SENNA 219 broader
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1982] CASSIINAE—SENNA 223 Pohl."
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1^^2] CASSIINAE—SENNA 225 between
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1982] CASSIINAE—SENNA 227 sometim
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1982] CASSIINAE—SENNA 229 sometim
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1982] CASSIINAE—SENNA 231 the hor
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1982] CASSIINAE—SENNA 233 paratyp
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1982] CASSIINAE—SENNA 237 tiary c
Page 249 and 250:
1982] CASSIINAE—SENNA 239 2.5-9(-
Page 251 and 252:
1982] CASSIINAE—SENNA 241 9. Pod
Page 253 and 254:
1982] CASSIINAE—SENNA 243 Cuba; d
Page 255 and 256:
1982] CASSIINAE—SENNA 245 the ste
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1982] CASSIINAE—SENNA 247 in outh
Page 259 and 260:
1982] CASSIINAE—SENNA 249 "In Bra
Page 261 and 262:
1982] CASSIINAE—SENNA 251 the lon
Page 263 and 264:
1982] CASSIINAE—SENNA 253 fainriy
Page 265 and 266:
1982] CASSIINAE—SENNA 255 is not
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1982] CASSIINAE—SENNA 257 70b. Se
Page 269 and 270:
1982] CASSIINAE—SENNA 259 racemes
Page 271 and 272:
1982] CASSUNAE—senna 261 other co
Page 273 and 274:
1982] CASSIINAE—SENNA 263 largely
Page 275 and 276:
Page 277 and 278:
1982] CASSIINAE—SENNA 267 sides t
Page 279 and 280:
1982] CASSIINAE—SENNA 269 found,
Page 281 and 282:
1982] CASSIINAE—SENNA 271 cordate
Page 283 and 284:
1982] CASSIINAE—SENNA 273 and hya
Page 285 and 286:
1982] CASSIINAE—SENNA 275 Key to
Page 287 and 288:
1982] CASSIINAE—SENNA 277 Peduncl
Page 289 and 290:
1982] CASSIINAE—SENNA 279 margine
Page 291 and 292:
1982] CASSIINAE—SENNA 281 Sandy b
Page 293 and 294:
1982] CASSIINAE—SENNA 283 cm, at
Page 295 and 296:
1982] CASSIINAE—SENNA 285 like th
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1982] CASSIINAE—SENNA 289 not con
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1982] CASSIINAE—SENNA 291 almost
Page 303 and 304:
1982] CASSIINAE—SENNA 293 filamen
Page 305 and 306:
1982] CASSIINAE—SENNA 295 2. Lfts
Page 307 and 308:
1982] CASSIINAE—SENNA 297 2. Fls
Page 309 and 310:
1982] CASSIINAE—SENNA 299 Main ca
Page 311 and 312:
Page 313 and 314:
1982] CASSIINAE—SENNA 303 a trifl
Page 315 and 316:
1982] CASSIINAE—SENNA 305 sentiaU
Page 317 and 318:
1982] CASSIINAE—SENNA 307 PUosulo
Page 319 and 320:
1982] CASSUNAE—senna 309 stamens,
Page 321 and 322:
1982] CASSIINAE—SENNA 311 of 3 ab
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1982] CASSIINAE—SENNA 315 Pubesce
Page 327 and 328:
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1982] CASSIINAE—SENNA 319 ovate,
Page 331 and 332:
Page 333 and 334:
1982] CASSIINAE—SENNA 323 buds su
Page 335 and 336:
1982] CASSIINAE—SENNA 325 orbata
Page 337 and 338:
1982] CASSIINAE—SENNA 327 glabrou
Page 339 and 340:
1982] CASSUNAE—senna 329 nation o
Page 341 and 342: 1982] CASSIINAE—SENNA 331 varying
Page 343 and 344: 1982] CASSIINAE—SENNA 333 seldom
Page 345 and 346: 1982] CASSIINAE—SENNA 335 to and
Page 347 and 348: 1982] CASSIINAE—SENNA 337 Charact
Page 349 and 350: 1982] CASSIINAE—SENNA 339 Key to
Page 351 and 352: 1982] CASSIINAE—SENNA 341 Fig. 26
Page 353 and 354: 1982] CASSIINAE—SENNA 343 Dry roc
Page 355 and 356: 1982] CASSIINAE—SENNA 345 llOg. S
Page 357 and 358: 1982] CASSIINAE—SENNA 347 exempli
Page 359 and 360: 1982] CASSIINAE—SENNA 349 17. Lft
Page 361 and 362: 1982] CASSIINAE—SENNA 351 (3-)4-6
Page 363 and 364: 1982] CASSUNAE—senna 353 a photog
Page 365 and 366: 1982] CASSUNAE—senna 355 the pedu
Page 367 and 368: 1982] CASSIINAE—SENNA 357 filifor
Page 369 and 370: 1982] CASSIINAE—SENNA 359 Stamens
Page 371 and 372: 1982] CASSIINAE—SENNA 361 Pods "o
Page 373 and 374: 1982] CASSIINAE—SENNA 363 obtuse
Page 375 and 376: 1982] CASSIINAE—SENNA 365 the vex
Page 377 and 378: 1982] CASSIINAE—SENNA 367 Racemes
Page 379 and 380: 1982] CASSIINAE—SENNA 369 Stipule
Page 381 and 382: 1982] CASSIINAE—SENNA 371 cm, the
Page 383 and 384: 1982] CASSIINAE—SENNA 373 Pod pen
Page 385 and 386: 1982] CASSIINAE—SENNA 375 Stipule
Page 387 and 388: 1982] CASSIINAE—SENNA 377 chilens
Page 389 and 390: 1982] CASSIINAE—SENNA 379 mm, the
Page 391: 1982] CASSIINAE—SENNA 381 found t
Page 395 and 396: 1982] CASSIINAE—SENNA 385 mostly
Page 397 and 398: 1982] CASSIINAE—SENNA 387 replace
Page 399 and 400: 1982] CASSIINAE—SENNA 389 eastern
Page 401 and 402: 1982] CASSIINAE—SENNA 391 h is ju
Page 403 and 404: 1982] CASSIINAE—SENNA 393 in sylv
Page 405 and 406: 1982] CASSIINAE—SENNA 395 Araguai
Page 407 and 408: 1982] CASSIINAE—SENNA 397 or almo
Page 409 and 410: 1982] CASSIINAE—SENNA 399 Pod ess
Page 411 and 412: 1982] CASSIINAE—SENNA 401 only ±
Page 413 and 414: 1982] CASSIINAE—SENNA 403 locaUy
Page 415 and 416: 1982] CASSIINAE—SENNA 405 COLUTEO
Page 417 and 418: 1982] CASSIINAE—SENNA 407 within
Page 419 and 420: 1982] CASSIINAE—SENNA 409 25. Pod
Page 421 and 422: 1982] CASSIINAE—SENNA 411 but the
Page 423 and 424: 1982] CASSIINAE—SENNA 413 interfo
Page 425 and 426: 1982] CASSIINAE—SENNA 415 cm; bra
Page 427 and 428: 1982] CASSIINAE—SENNA 417 -35) mm
Page 429 and 430: 1982] CASSUNAE—senna 419 mexicana
Page 431 and 432: 1982] CASSIINAE—SENNA 421 places,
Page 433 and 434: 1982] CASSIINAE—SENNA 423 Key to
Page 435 and 436: 1982] CASSIINAE—SENNA 425 137d. S
Page 437 and 438: 1982] CASSIINAE—SENNA 427 across
Page 439 and 440: 1982] CASSIINAE—SENNA 429 6. Vena
Page 441 and 442: 1982] CASSIINAE—SENNA 431 on the
Page 443 and 444:
1982] CASSIINAE—SENNA 433 var. gl
Page 445 and 446:
1982] CASSIINAE—SENNA 435 Caledon
Page 447 and 448:
1982] CASSIINAE—SENNA 437 occiden
Page 449 and 450:
1982] CASSIINAE—SENNA 439 piano-c
Page 451 and 452:
1982] CASSIINAE—SENNA 441 Bot. 24
Page 453 and 454:
1982] CASSIINAE—SENNA 443 valves
Page 455 and 456:
1982] CASSUNAE—senna 445 scarcely
Page 457 and 458:
1982] CASSIINAE—SENNA 447 Cassia
Page 459 and 460:
1982] CASSIINAE—SENNA 449 Sesse &
Page 461 and 462:
1982] CASSIINAE—SENNA 451 Neg. 24
Page 463 and 464:
1982] CASSUNAE—senna 453 tierra t
Page 465:
^m\m'^Y%^ws:m.yitmxmfsmsiiimmammKmm
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