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Cassiinae pt 1 NY-Botanical_gardens_Vol. 35_1 - Copy.pdf - Antbase

Cassiinae pt 1 NY-Botanical_gardens_Vol. 35_1 - Copy.pdf - Antbase

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64 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [VOL. <strong>35</strong><br />

SENNA [K. Bauhin] P. MiUer<br />

Senna [K. Bauhin] P. MiUer, Gard. Diet., Abr. ed. 4, vol. 3. 1754, sensu magnopere<br />

ampUat. Irwin & Barneby in PolhiU & Raven, 1981, p. 105.—<br />

Generitypus: ''Senna alexandrina sive foliis acutis C[aspar] B[auhin]<br />

P[inax 397]" = Senna alexandrina P. MiUer, 176S.—Cassia sect. Sen­<br />

na [Tournefort] DeCandoUe ex Colladon, Hist. Casses 92. 1816. Cas­<br />

sia subgen. Senna Bentham in Martius, Fl. Bras. 15(2): 96. 1870;<br />

Trans. Linn. Soc. London 27: 513, 518. 1871.<br />

Senna sensu Gaertner, Fruct. sem. pl. 2: 312, t. 146, fig. 4. 1791; Link, Handbuch 2: 139. 1831;<br />

Roxburgh, Fl. Ind. ed. Carey, 2: 339. 1832.<br />

Cassia sect. Senna sensu Bentham in Bentham & Hooker, Gen. Pl. 1(2): 139. 1865.<br />

General characters of subtribe <strong>Cassiinae</strong>, with these parricular ones: pedicels<br />

ebracteolate (exce<strong>pt</strong> S. peralteana and S. paradictyon); androecium (Fig. 10, 11)<br />

zygomorphic, funcrionaUy 4-10-merous, the stamens ± graduated in length from<br />

ad- to abaxial side of fl, the 3 adaxial stamens (exce<strong>pt</strong> in Palaeotropic sect. Psi­<br />

lorhegma) staminodal, the (6-)7 fertUe anthers basifixed, usuaUy modified into<br />

heteromorphic sets of 4 median and (2-)3 abaxial (these lacking in S. hayesiana),<br />

all beakless or variably beaked or appendaged at apex, where dehiscent by 2<br />

separate or confluent pores or short shts, the thecae either glabrous or pubescent<br />

but not ciholate along the lateral (closed) sutures; pod either dehiscent or indehiscent,<br />

if the former inerdy so through both sutures, the valves not elasticaUy<br />

coihng, when indehiscent either breaking up into 1-seeded achenehke segments<br />

or the seeds released only by rotting of the valves; seed-funicle filiform; seedcoat<br />

commonly but not always areolate, never serially pitted.—Trees, shrubs,<br />

herbs, some monocarpic, the roots often blackish, lacking bacterial nodules;<br />

extrafloral nectaries (Fig. 9), when present, ovoid, globose, claviform or phalloid,<br />

excreting nectar from convex surface (plane in petiolar gland of S. kuhlmannii);<br />

foliage often foetid; lfts 1-many pairs or lvs rarely (exce<strong>pt</strong> for transient eophylls)<br />

absent or (in AustraUa) phyllodal; inflorescence of 1-many-fld racemes, these<br />

axillary from hornotinous stems or sometimes cauliforous, then either sohtary or<br />

paniculate; hypanthium sohd, shallowly bowl-shaped or slenderly vase-shaped,<br />

often not externally differentiated from pedicel; calyx-lobes either equal or<br />

strongly graduated, the inner then longer, mostly obtuse; petals (in America al­<br />

ways) yeUow, rarely white, drying either yeUow (brown) or whitish dark-veined,<br />

the corolla either zygomorphic or irregular, one abaxial petal then highly modified<br />

either as a flag or as an androecial shield, the vexUlar petal almost always interior<br />

in praeflorarion; pistU either centric or enantiostylous; ovules 5-266; stigma punc­<br />

tiform or dilated and cupular, usually cUiolate; pod (Fig. 12-14) highly variable<br />

in length, texture and compression, primirively piano-compressed, becoming turgid,<br />

angulate, terete, or rarely either its valves or its sutures winged lengthwise,<br />

the valves sometimes crested or otherwise elevated over seeds, the dry or pulpfilled<br />

cavity continuous or commonly transversely se<strong>pt</strong>ate, the valves commonly<br />

conrinu ous, sometimes breaking up into panels or into indehiscent achenehke<br />

segments; seeds free or (Madagascan Senna perrieri Ghesqu.) adherent to the<br />

endocarp, mostly 1-, less often 2-, exce<strong>pt</strong>ionally 4-seriate, either transverse or<br />

basipetal in orientation, lying broadside either to the valves or to the interseminal<br />

se<strong>pt</strong>a, their testa naked or wax-coated (crackled when ripe), smooth or irregularly<br />

pitted, areolate or not; 2n = 22, 23, 26, 28.—Spp. ±260, about "^/s's American,<br />

these referred below to <strong>35</strong> series in 6 sections, the whole genus circumtropical<br />

but extending into warm temperate, temperate desert-grassland and exceprionaUy

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