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Aretz et al_2011.pdf - ORBi - Université de Liège

Aretz et al_2011.pdf - ORBi - Université de Liège

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Kölner Forum Geol. P<strong>al</strong>äont., 19 (2011)<br />

M. ARETZ, S. DELCULÉE, J. DENAYER & E. POTY (Eds.)<br />

Abstracts, 11th Symposium on Fossil Cnidaria and Sponges, <strong>Liège</strong>, August 19-29, 2011<br />

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98<br />

Infraspecific variation in actinastreid cor<strong>al</strong>s (Scleractinia; Jurassic-<br />

Cr<strong>et</strong>aceous) and implications for the species concept<br />

Hannes LÖSER<br />

Universidad Nacion<strong>al</strong> Autónoma <strong>de</strong> México, Instituto <strong>de</strong> Geología, Estación Region<strong>al</strong> <strong>de</strong>l Noroeste, Col.<br />

Los Arcos, L.D.Colosio s/n, 83200 Hermosillo, Sonora, Mexico; loeser@p<strong>al</strong>eotax.<strong>de</strong><br />

Cor<strong>al</strong>s of the family Actinastraeidae are extremely common in the Late Jurassic and Early Cr<strong>et</strong>aceous<br />

and various genera have a high number of species. Species separation is gener<strong>al</strong>ly based on c<strong>al</strong>icular<br />

dimensions (diam<strong>et</strong>er, distance, amount per given area), sept<strong>al</strong> symm<strong>et</strong>ry, and sept<strong>al</strong> number. To obtain<br />

b<strong>et</strong>ter insight into intraspecific variation and results for species separation, systematic measurements of the<br />

cor<strong>al</strong>s were taken and statistic<strong>al</strong>ly an<strong>al</strong>ysed. As a preliminary study, ten type specimens were selected for<br />

an<strong>al</strong>ysis. In large thin sections a large number of c<strong>al</strong>ices (up to 200) were measured, including their<br />

diam<strong>et</strong>er, distance and the thickness of the w<strong>al</strong>l and coenosteum. For <strong>al</strong>l v<strong>al</strong>ues, the arithm<strong>et</strong>ic mean,<br />

standard <strong>de</strong>viation and the coefficient of variation were c<strong>al</strong>culated. It was observed, that the c<strong>al</strong>icular<br />

diam<strong>et</strong>er is the character with the lowest variation whereas the distance of the c<strong>al</strong>icular centres, the<br />

thickness of the w<strong>al</strong>l, and the number of c<strong>al</strong>ices per a given area show a much higher variation and are<br />

therefore not suitable for distinguishing samples within a population or species of different faunas. It was<br />

found that about 70% of <strong>al</strong>l v<strong>al</strong>ues of the lumen diam<strong>et</strong>er f<strong>al</strong>l in the first interv<strong>al</strong> (range of the arithm<strong>et</strong>ic<br />

mean ± standard <strong>de</strong>viation). Hence, the first interv<strong>al</strong> is a good representation for most types of measured<br />

v<strong>al</strong>ues in fossil cor<strong>al</strong>s.<br />

To obtain a representative v<strong>al</strong>ue for the arithm<strong>et</strong>ic mean, only a low number of measurements is<br />

required (20-30), but in or<strong>de</strong>r to obtain representative v<strong>al</strong>ues for the standard <strong>de</strong>viation, the coefficient of<br />

variation and the first interv<strong>al</strong> more v<strong>al</strong>ues are necessary (> 50). The results are compared to tradition<strong>al</strong><br />

m<strong>et</strong>hods by remeasuring published materi<strong>al</strong>. It is conclu<strong>de</strong>d that the application of systematic measuring<br />

should be exten<strong>de</strong>d to other species rich cor<strong>al</strong> genera.<br />

Fig. 1: Screen shot of the program used to measure the dimensions of the cor<strong>al</strong>s (P<strong>al</strong>eoTax/Measure).

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