Aretz et al_2011.pdf - ORBi - Université de Liège
Aretz et al_2011.pdf - ORBi - Université de Liège
Aretz et al_2011.pdf - ORBi - Université de Liège
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Kölner Forum Geol. P<strong>al</strong>äont., 19 (2011)<br />
M. ARETZ, S. DELCULÉE, J. DENAYER & E. POTY (Eds.)<br />
Abstracts, 11th Symposium on Fossil Cnidaria and Sponges, <strong>Liège</strong>, August 19-29, 2011<br />
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98<br />
Infraspecific variation in actinastreid cor<strong>al</strong>s (Scleractinia; Jurassic-<br />
Cr<strong>et</strong>aceous) and implications for the species concept<br />
Hannes LÖSER<br />
Universidad Nacion<strong>al</strong> Autónoma <strong>de</strong> México, Instituto <strong>de</strong> Geología, Estación Region<strong>al</strong> <strong>de</strong>l Noroeste, Col.<br />
Los Arcos, L.D.Colosio s/n, 83200 Hermosillo, Sonora, Mexico; loeser@p<strong>al</strong>eotax.<strong>de</strong><br />
Cor<strong>al</strong>s of the family Actinastraeidae are extremely common in the Late Jurassic and Early Cr<strong>et</strong>aceous<br />
and various genera have a high number of species. Species separation is gener<strong>al</strong>ly based on c<strong>al</strong>icular<br />
dimensions (diam<strong>et</strong>er, distance, amount per given area), sept<strong>al</strong> symm<strong>et</strong>ry, and sept<strong>al</strong> number. To obtain<br />
b<strong>et</strong>ter insight into intraspecific variation and results for species separation, systematic measurements of the<br />
cor<strong>al</strong>s were taken and statistic<strong>al</strong>ly an<strong>al</strong>ysed. As a preliminary study, ten type specimens were selected for<br />
an<strong>al</strong>ysis. In large thin sections a large number of c<strong>al</strong>ices (up to 200) were measured, including their<br />
diam<strong>et</strong>er, distance and the thickness of the w<strong>al</strong>l and coenosteum. For <strong>al</strong>l v<strong>al</strong>ues, the arithm<strong>et</strong>ic mean,<br />
standard <strong>de</strong>viation and the coefficient of variation were c<strong>al</strong>culated. It was observed, that the c<strong>al</strong>icular<br />
diam<strong>et</strong>er is the character with the lowest variation whereas the distance of the c<strong>al</strong>icular centres, the<br />
thickness of the w<strong>al</strong>l, and the number of c<strong>al</strong>ices per a given area show a much higher variation and are<br />
therefore not suitable for distinguishing samples within a population or species of different faunas. It was<br />
found that about 70% of <strong>al</strong>l v<strong>al</strong>ues of the lumen diam<strong>et</strong>er f<strong>al</strong>l in the first interv<strong>al</strong> (range of the arithm<strong>et</strong>ic<br />
mean ± standard <strong>de</strong>viation). Hence, the first interv<strong>al</strong> is a good representation for most types of measured<br />
v<strong>al</strong>ues in fossil cor<strong>al</strong>s.<br />
To obtain a representative v<strong>al</strong>ue for the arithm<strong>et</strong>ic mean, only a low number of measurements is<br />
required (20-30), but in or<strong>de</strong>r to obtain representative v<strong>al</strong>ues for the standard <strong>de</strong>viation, the coefficient of<br />
variation and the first interv<strong>al</strong> more v<strong>al</strong>ues are necessary (> 50). The results are compared to tradition<strong>al</strong><br />
m<strong>et</strong>hods by remeasuring published materi<strong>al</strong>. It is conclu<strong>de</strong>d that the application of systematic measuring<br />
should be exten<strong>de</strong>d to other species rich cor<strong>al</strong> genera.<br />
Fig. 1: Screen shot of the program used to measure the dimensions of the cor<strong>al</strong>s (P<strong>al</strong>eoTax/Measure).