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Beetles Identification Guide

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Economic importance: The species is a minor pest in Canada, where it has been<br />

found in flour and feed mills, warehouses, dwellings, and occasionally in grain<br />

samples from granaries. In other regions of the world, it is considered to be a<br />

major or potentially serious pest of stored products, particularly seeds, cereals,<br />

legumes, nuts, and other high-protein food.<br />

Selected references<br />

Archer, T.L.; Strong, R.G. 1975. Comparative studies on the biologies of six<br />

species of Trogoderma: T. glabrum. Ann. Entomol. Soc. Am. 68:105—114.<br />

Azab, A.K., Tawfik, M.F.S.; Abouzeid, N.A. 1973. The biology of Dermestes<br />

maculatus De Geer (Coleoptera: Dermestidae). Bull. Soc. Entomol. Egypte<br />

56:1—14.<br />

Barak, A.V.; Burkholder, W.E. 1977. Studies on the biology of Attagenus<br />

elongatulus Casey (Coleoptera: Dermestidae) and the effects of larval<br />

crowding on pupation and life cycle. J. Stored Prod. Res. 13:169—175.<br />

Beal, R.S. 1954. Biology and taxonomy of the Nearctic species of Trogoderma<br />

(Coleoptera: Dermestidae). Univ. Calif. Publ. Entomol. 10:35—102.<br />

Beal, R.S. 1956. Synopsis of the economic species of Trogoderma occurring in<br />

the United States with description of a new species (Coleoptera:<br />

Dermestidae). Ann. Entomol. Soc. Am. 49:559—566.<br />

Beal, R.S. 1970. A taxonomic and biological study of species of Attagenini<br />

(Coleoptera: Dermestidae) in the United States and Canada. Entomol.<br />

Am. 45:141—235.<br />

Burges, H.D. 1960. Studies on the dermestid beetle Trogoderma granarium<br />

Everts—IV. Feeding, growth, and respiration with particular reference to<br />

diapause larvae. J. Insect Physiol. 5:317—334.<br />

Coombs, C.W. 1981. The development, fecundity and longevity of Dermestes<br />

ater DeGeer (Coleoptera: Dermestidae). J. Stored Prod. Res. 17:31—36.<br />

Franciscolo, M.E. 1975. Sulla posizione sistematica di Thylodrias contractus<br />

Motschulsky 1839 (Coleoptera, Series Bostrychiformia). Boll. Soc. Entomol.<br />

Ital. 107:142—146.<br />

Green, M. 1979. The identification of Trogoderma variabile Ballion, T. inclusum<br />

LeConte and T. granarium Everts (Coleoptera: Dermestidae) using<br />

characters provided by their genitalia. Entomol. Gaz. 30:199—204.<br />

Greenwald, M. 1941. Studies on the biology of four common carpet beetles.<br />

Part II. The old fashioned carpet beetle (Anthrenus scrophulariae L.). Agric.<br />

Exp. Stn. Univ. Cornell Mem. 240:58—75.<br />

Griswold, G.H. 1941. Studies on the biology of four common carpet beetles.<br />

Part I. The black carpet beetle (Attagenus piceus Oliv.), the varied carpet<br />

beetle (Anthrenus verbasci L.), and the furniture carpet beetle (Anthrenus<br />

vorax Waterh.). Agric. Exp. Stn. Univ. Cornell Mem. 240:1—57.<br />

Halstead, D.G.H. 1981. Taxonomic notes on some Attagenus spp. associated<br />

with stored products, including a new black species from Africa (Coleoptera:<br />

121

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