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Lesson 3: Species in the environmental complex

Lesson 3: Species in the environmental complex

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Why do plants grow where <strong>the</strong>y do?<br />

The holocoenotic<br />

<strong>environmental</strong><br />

<strong>complex</strong><br />

• A <strong>complex</strong>, <strong>in</strong>dvisible<br />

whole system<br />

consist<strong>in</strong>g of <strong>the</strong> plant<br />

and its multitude of<br />

<strong>environmental</strong><br />

<strong>in</strong>fluences.<br />

• Holocoenosis =<br />

ecosystem?<br />

• Compare to Tansley s<br />

concept of <strong>the</strong><br />

“ecosystem”, which<br />

also <strong>in</strong>cluded time and<br />

change.<br />

• Bill<strong>in</strong>g s concept is<br />

<strong>the</strong>refore an<br />

ecosystem at a<br />

moment <strong>in</strong> time.<br />

W.D. Bill<strong>in</strong>gs, 1952. The <strong>environmental</strong> <strong>complex</strong> <strong>in</strong> relation to plant growth and<br />

distribution. Quaterly Review of Biology 27: 251-265.

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