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A land manager's guide to conserving habitat for forest birds in ...

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Elderberry — Pho<strong>to</strong>: Robert McCaw<br />

like the Wood Thrush will respond positively <strong>to</strong> the structural<br />

changes associated with low <strong>in</strong>tensity selection system harvest<strong>in</strong>g<br />

and have higher nest<strong>in</strong>g success <strong>in</strong> these sites. However, this<br />

benefit does not occur until at least five years after harvest, but<br />

can persist <strong>for</strong> a number of years (5–10 years).<br />

Predation and parasitism are not the only changes associated<br />

with harvest<strong>in</strong>g that can affect avian productivity. Reductions <strong>in</strong><br />

food availability can strongly <strong>in</strong>fluence a parent’s ability <strong>to</strong> feed<br />

its nestl<strong>in</strong>gs. This can result <strong>in</strong> fewer young or less healthy young<br />

produced by the pair of <strong>birds</strong>. Research shows both Northern<br />

Card<strong>in</strong>als and Rose-breasted Grosbeaks fledge fewer young per<br />

nest <strong>in</strong> diameter-limit sites than uncut sites. This may be due <strong>to</strong><br />

lower food availability, <strong>in</strong>creased parasitism (as cow<strong>birds</strong> typically<br />

remove a host egg), or both. Virtually all song<strong>birds</strong> eat <strong>in</strong>sects<br />

and other <strong>in</strong>vertebrates dur<strong>in</strong>g the breed<strong>in</strong>g season. When you<br />

open up the canopy by harvest<strong>in</strong>g, a <strong>for</strong>est becomes sunnier,<br />

hotter, and drier. This can reduce the biomass of <strong>in</strong>sects that are<br />

vulnerable <strong>to</strong> desiccation. The changes <strong>in</strong> micro-climate can be<br />

extreme as the <strong>in</strong>tensity of harvest <strong>in</strong>creases, and translate <strong>in</strong><strong>to</strong><br />

larger changes <strong>in</strong> <strong>for</strong>est structure and <strong>in</strong>sect communities.<br />

Reductions <strong>in</strong> the density of mature trees can result <strong>in</strong> less bark<br />

and canopy dwell<strong>in</strong>g <strong>in</strong>sects available <strong>for</strong> canopy and bark<br />

feeders. In selection system harvests where you reta<strong>in</strong> most of the<br />

mature canopy and the abiotic environment changes little, <strong>in</strong>sect<br />

communities will be more similar <strong>to</strong> uncut <strong>for</strong>ests. Fly<strong>in</strong>g <strong>in</strong>sects,<br />

on the other hand, can respond positively <strong>to</strong> abiotic changes<br />

created by canopy gaps, and thus group cuts can create ideal<br />

<strong>for</strong>ag<strong>in</strong>g <strong>habitat</strong> <strong>for</strong> aerial <strong>in</strong>sectivores, such as flycatchers. As<br />

<strong>in</strong>sects are not the only source of food, changes <strong>in</strong> <strong>for</strong>est structure<br />

and composition may alter availability of fruits and seeds.<br />

Growth and regeneration of mid-<strong>to</strong>lerant tree species like red<br />

and white oak, bitternut hickory and black cherry through<br />

shelterwood or group selection can <strong>in</strong>crease the availability of<br />

tree seeds (also known as mast). Similarly, gaps will often<br />

stimulate shrub growth, creat<strong>in</strong>g new sources of food, like<br />

raspberry or elderberry. Thus harvest<strong>in</strong>g can have positive or<br />

negative effects on food availability depend<strong>in</strong>g on species-specific<br />

<strong>for</strong>ag<strong>in</strong>g preferences and the system employed.<br />

Survival of juveniles after they leave or fledge from the nest is<br />

another key component <strong>in</strong>fluenc<strong>in</strong>g avian productivity. Dur<strong>in</strong>g<br />

the post-breed<strong>in</strong>g period, adults and juveniles of mature <strong>for</strong>est<br />

bird species start <strong>to</strong> make use of early successional <strong>habitat</strong>s,<br />

where an open canopy creates new growth <strong>in</strong> the <strong>for</strong>m of a dense<br />

shrub layer. These <strong>habitat</strong>s offer greater protection from<br />

preda<strong>to</strong>rs, and <strong>for</strong> some species, high food sources. For species<br />

like Hooded Warbler, heavy diameter-limit cut sites may actually<br />

have higher juvenile survival rates than s<strong>in</strong>gle tree selection or<br />

unharvested sites. While <strong>for</strong> other species like Rose-breasted<br />

Grosbeaks, harvest treatment has little impact on fledgl<strong>in</strong>g<br />

survival parameters. Thus, early successional <strong>habitat</strong>s can provide<br />

valuable <strong>habitat</strong> <strong>to</strong> juvenile <strong>birds</strong> of some species but do very<br />

little <strong>for</strong> another.<br />

In summary, the <strong>in</strong>tensity of the harvest strongly <strong>in</strong>fluences<br />

the extent <strong>to</strong> which harvest<strong>in</strong>g affects bird communities. Evenaged<br />

management creates very different breed<strong>in</strong>g conditions and<br />

different bird communities, than uneven-aged management. In a<br />

cont<strong>in</strong>uously <strong>for</strong>ested <strong>land</strong>scape (like the boreal <strong>for</strong>est region),<br />

even-aged management m<strong>in</strong>imizes many of the dramatic changes<br />

associated with harvest<strong>in</strong>g, s<strong>in</strong>ce the bird species are adapted <strong>to</strong><br />

<strong>in</strong>frequent large-scale disturbances. Loss of canopy cover and<br />

Rose-breasted Grosbeak fledgl<strong>in</strong>gs — Pho<strong>to</strong>: Brad Woodworth<br />

Harvest<strong>in</strong>g Effects on Birds 55

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