Part 7. JUNCACEAE 2: Juncus - Species Plantarum Programme

8
SPECIES PLANTARUM — FLORA OF THE WORLD (2002)
10. GREAT BRITAIN: England, Norfolk, Burnham Overy, A.C.Jermy & G.Simmons 535 (BM).
12. FRANCE: Bouches-du-Rhône, Les Stes Maries, S.Snogerup 360 (LD). CORSE: Ajaccio, H.Roessler
3557 (M). 13. GREECE: Kerkyra, S.Snogerup 23608 (LD). 20. ALGERIA: between Boufakir and Kolea,
P.H.Davis 59506 (E, BM). EGYPT: El Dikheila, M.N.El Hadidi s.n. (LD). 34. IRAN: Nov-Char, Sabeti
111 (W). JORDAN: Azraq, V.Täckholm 9088 (CAI, K). TURKEY: Istanbul, Catalca, Tutel & Delice
32935 (ISTF). 50. NEW SOUTH WALES: Clyde R., Nelligen, E.F.Constable 43125 (UC). 51. NORTH
ISLAND: Papuke turnoff at head of Whangaroa inlet, R.Melville 5346, 12 Nov 1961 (NSW).
1b. Juncus acutus subsp. leopoldii (Parl.) Snogerup, Bot. Not. 130: 187 (1978)
Juncus leopoldii Parl., Giorn. Bot. Ital. 2(1): 324 (1846); Juncus acutus var. leopoldii (Parl.) Buchenau,
Monogr. Junc. Cap 421 (1875) [Abh. Naturwiss. Ver. Bremen 4: 421 (1875)]. T: South Africa, Sommerset,
Stellenbosch, C.F.Ecklon & C.Zeyher 4308; lecto: FI, fide S.Snogerup, Willdenowia 23: 37 (1993); iso: S, W.
Juncus acutus L. var. sphaerocarpus Engelm., in J.Wheeler, Rep. U.S. Geol. Survey, Bot. 6: 376 (1878);
Juncus robustus S.Watson, Proc. Amer. Acad. Arts 14: 302 (1879). T: California, S. Barbara [not found].
Juncus acutus L. f. xanthosus Jeps., Fl. Calif. 1: 249 (1921). T: California, NW of Indio, Thousand Palms
Cañon, W.L.Jepson 6041; holo: UC.
Illustration: S.Snogerup, Willdenowia 23: 35, fig. 2, J–O (1993).
Map: S.Snogerup, Willdenowia 23: 36, 68, fig. 22 (1993).
Stem (50–) 80–150 (–200) cm tall, usually 1.5–2.5 (–6) mm in diam.; leaves 40–100 cm long,
1.5–3 mm in diam.; sheaths wide, up to 25 cm long. Inflorescence 5–30 (–40) × 2–6 (–8) cm,
lax or rarely congested, with (10–) 35–100 (–250) heads; heads often densely aggregated into
±large groups. Tepals 2.2–3.7 mm long; outer tepals inserted slightly below the inner, oblong
with broad hyaline margins; inner tepals cymbiform with small distal auricles; mucro up to
0.5 mm long. Anthers 1.4–1.8 mm long, (2–) 2.5–4 times as long as filaments. Style 0.6–1.3 mm
long; stigmas 1.5–2.5 mm long. Capsule (3–) 3.5–5 mm long, obovoid, usually castaneous to
dark or rarely light brown, basally lighter, trilocular or when ripe triseptate; apex usually
obtuse; mucro 0.2–0.6 mm. Seeds 0.7–0.8 (–0.9) mm long, or 1.1–1.4 mm long including
appendages. 2n = 48, fide S.Snogerup, Willdenowia 23: 39 (1993). Fig. 38.
Large disjunct distribution range: Atlantic Islands, NW African Coast and Iberian Peninsula,
South Africa, C temperate South America, W coast of Mexico, California and Nevada,
Bermuda. 12: POR, SPA. 20: EGY, LBY, MOR. 21: CNY, CVI, MDR. 27: CPP, LES, NAM,
NAT, OFS. 28: ASC. 76: ARI, CAL, NEV. 79: MXN. 81: BER. 84: BZS. 85: AGE, AGS,
AGW, CLN, CLS, JNF, URU. Coastal sandy habitats, salt marshes and inland saline or
marshy habitats, from sea level to 1400 m (in South Africa). Map 175 [the map includes
intermediates between the subspecies in the Azores and Iberian Peninsula].
12. PORTUGAL: Estremadura, Sacavém, Rio Trancão, Foules & Rainha 15381 (S). SPAIN: La Coruña,
2 Oct 1852, J.M.C.Lange (C, GB). 20. MOROCCO: Tanger, H.Lindberg 1532 (H). 21. CANARY IS.:
Gran Canaria, E.Wängsjö 231 (LD). 27. CAPE PROVINCE: Laingsburg, R.H.Compton 6761 (BOL).
76. CALIFORNIA: Los Angeles, S.Snogerup & B.Snogerup 3850 (LD). 85. ARGENTINA NORTHWEST:
Salta, Chicoana, La Maroma, Río Arias, Cabra Corral Dam, 1030 m, L.J.Novara 10577 (MCNS).
ARGENTINA NORTHEAST: Buenos Aires, Laguna Sauce Grande, R.Santesson 97 (S). CHILE: Antofagasta,
Taltal, Johnston 5126 (K).
Rare intermediates between the subspecies are known from SW Europe, Libya and Egypt,
and Azores [e.g., Egypt, Ain Murun, Ascherson 513 (CAI, G, K, M, W, WU)]. Occurrence in
NW Africa, Spain and Portugal may be a result of a relatively recent introduction.
2. Juncus littoralis C.A.Mey., Verz. Pfl. Casp. Meer. 34 (1831)
Juncus acutus L. var. littoralis (C.A.Mey.) Trautv., Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 480 (1831);
Juncus acuto-maritimus E.Mey., in C.F. von Ledebour, Fl. Ross. 4: 234 (1853), nom. illeg.; Juncus acutus
subsp. littoralis (C.A.Mey.) Feinbrun, Fl. Palaestina 4: 143 (1986). T: Insula Sara, 26 Jun 1830, C.A.Meyer
138; lecto: LE, fide S.Snogerup, in K.H.Rechinger, Fl. Iranica 75: 6 (1971); iso: G, G-BOIS, W.
Juncus tommasinii Parl., Fl. Ital. 2: 315 (1852); Juncus acutus L. subsp. tommasinii (Parl.) Arcang., Comp. Fl.
Ital. 715 (1882); Juncus acutus L. var. tommasinii (Parl.) Buchenau, Bot. Jahrb. Syst. 12: 250 (1890); Juncus
acutus proles tommasinii (Parl.) Rouy, Fl. France 13: 230 (1912). T: [Italy] 'Nei paludi presso Monfalcone,
Grado', M.G.S.Tommasini, lecto: FI, fide S.Snogerup, Willdenowia 23: 40 (1993).