ERDC/EL TR-12-5 10mineral soils (Erickson <strong>and</strong> Puttock 2006). The principal wetl<strong>and</strong> types in<strong>the</strong> region are described briefly below.Depressional wetl<strong>and</strong>s occur on coastal plains behind beach or riverberms, <strong>and</strong> in historic fish ponds that have filled with sediment. Theirmain water source is groundwater, which can be fresh in <strong>the</strong> wet season tohypersaline in <strong>the</strong> dry season <strong>and</strong> may be influenced by storm surges. In<strong>the</strong> main <strong>Hawaii</strong>an Isl<strong>and</strong>s, <strong>the</strong>se wetl<strong>and</strong>s typically are found on mudflatsor in s<strong>and</strong>y areas, <strong>and</strong> support woody vegetation, such as saltwort (Batismaritima) <strong>and</strong> mangroves (Figure 3). In <strong>the</strong> Marianas <strong>and</strong> o<strong>the</strong>r <strong>Pacific</strong>isl<strong>and</strong>s, <strong>the</strong>y can also be forested with Nypa palms, Barringtonia racemosa,or o<strong>the</strong>r species. Examples <strong>of</strong> <strong>the</strong>se wetl<strong>and</strong>s can be found on <strong>the</strong>south shore <strong>of</strong> Moloka‘i, at Nukoli‘i on Kaua‘i, in <strong>the</strong> Kawainui Marsh <strong>and</strong>James Campbell National Wildlife Refuge on O‘ahu, <strong>and</strong> in <strong>the</strong> Kīheiwetl<strong>and</strong>s on Maui.Figure 3. Coastal wetl<strong>and</strong> on Kaua‘i dominated by Batis maritima <strong>and</strong> mangroves.Photo courtesy <strong>of</strong> <strong>the</strong> U.S. Fish <strong>and</strong> Wildlife Service.Sloped marshl<strong>and</strong>s or fern wetl<strong>and</strong>s obtain <strong>the</strong>ir hydrology from shallowgroundwater that may rise to <strong>the</strong> surface infrequently. They occur mainlyin volcanic areas on slopes greater than 10 percent, particularly in <strong>the</strong>
ERDC/EL TR-12-5 11Mariana Isl<strong>and</strong>s. They typically have mineral soils <strong>and</strong> support mainlyherbaceous vegetation (e.g., Nephrolepis ferns, Phragmites karka, <strong>and</strong>sedges), with scattered P<strong>and</strong>anus tectorius trees. Examples include <strong>the</strong>Shell wetl<strong>and</strong>s <strong>and</strong> Nimitz Hill marshl<strong>and</strong>s on Guam <strong>and</strong> <strong>the</strong> upper WaipāValley wetl<strong>and</strong>s on Kaua‘i.Hanging bogs occur in areas <strong>of</strong> abundant rainfall on steep volcanic slopes<strong>of</strong> 25 percent or more. Their hydrology is driven by groundwater perchedover an aquitard <strong>of</strong> ironstone sheath. The soils are <strong>of</strong>ten organic (e.g.,peats, mucks) <strong>and</strong> <strong>the</strong> vegetation is mainly herbaceous with stunted trees<strong>and</strong> shrubs (e.g., Metrosideros polymorpha). Examples can be found in<strong>the</strong> Waipā Valley <strong>of</strong> Kaua‘i <strong>and</strong> on high-elevation slopes in <strong>the</strong> Ko‘olauMountains on O‘ahu.Montane bogs are found mainly in volcanic areas that have flat to rollingtopography, abundant precipitation (up to 445 in. [11,300 mm] per year),<strong>and</strong> a high water table <strong>and</strong>/or surface flow. They are found in <strong>the</strong> AlakaiSwamp on Kaua‘i (Figure 4), on Mount Ka‘ala on O‘ahu, in <strong>the</strong> KamakouPreserve on Moloka‘i, on Mount ‘Eke <strong>and</strong> Hāna Ranch on Maui, <strong>and</strong> in <strong>the</strong>Mauna Loa bogs on Hawai‘i. Montane bogs are dominated by nativeherbaceous, fern, shrub, <strong>and</strong> stunted forest species.Forested wetl<strong>and</strong>s are found at middle to higher elevations on all <strong>of</strong> <strong>the</strong>larger <strong>Hawaii</strong>an Isl<strong>and</strong>s with <strong>the</strong> exception <strong>of</strong> Kaho‘olawe <strong>and</strong> Ni‘ihau. Theyoccupy slopes <strong>and</strong> depressions where <strong>the</strong> water table is <strong>of</strong>ten near <strong>the</strong>surface or where rainwater perches above a shallow aquitard or bedrock. In<strong>the</strong> Puna <strong>and</strong> South Hilo Districts on <strong>the</strong> Isl<strong>and</strong> <strong>of</strong> Hawai‘i, for example,many small wetl<strong>and</strong>s exist in scattered depressions created by cracks, folds,<strong>and</strong> undulating flow patterns in pāhoehoe lava flows (Wakeley et al. 1996).Riverine <strong>and</strong> open-water fringe wetl<strong>and</strong>s are present throughout <strong>the</strong> region.Their hydrology is derived mainly from <strong>the</strong> adjacent stream, lake, or pond.They are generally fresh water but may be brackish to saline at <strong>the</strong>irconfluence with marine waters. The vegetation <strong>of</strong> fringe wetl<strong>and</strong>s rangesfrom trees <strong>and</strong> shrubs (e.g., Rhizophora mangle, Xylocarpus moluccensis,<strong>and</strong> o<strong>the</strong>r mangroves; hau [Hibiscus tiliaceus]; <strong>and</strong> java plum [Syzygiumcumini = Eugenia cumini]) to herbaceous plants (e.g., para grass [Urochloamutica], sawgrass [Cladium mariscus ssp. jamaicense = C. jamaicense],<strong>and</strong> seashore paspalum [Paspalum vaginatum]). Throughout <strong>the</strong> region,this wetl<strong>and</strong> type represents <strong>the</strong> greatest acreage <strong>of</strong> remaining low-elevation