M.-S. Fletcher, P.I. Moreno / Quaternary International 253 (2012) 32e46 39than w25 km east of Lago Guanaco prior to AD 1881 (Dixie, 1881),imply<strong>in</strong>g that evaporation does not limit forest development <strong>in</strong> <strong>the</strong>vic<strong>in</strong>ity of Lago Guanaco today, and that precipitation exerts <strong>the</strong>dom<strong>in</strong>ant control over vegetation <strong>in</strong> that area. Despite this, it ispossible that, given <strong>the</strong> location of <strong>the</strong> site east of <strong>the</strong> orographicdivide, under vary<strong>in</strong>g westerly w<strong>in</strong>d regimes, <strong>the</strong> tension zonebetween evaporation and precipitation dom<strong>in</strong>ance over vegetationshifts east or west of its present location to <strong>the</strong> east of LagoGuanaco.Sediments retrieved from <strong>the</strong> Gran Campo-2 site (52 48 0 37 00 S,73 55 0 46 0 W; Fesq-Mart<strong>in</strong> et al., 2004), a m<strong>in</strong>erotropic mire locatedwest of <strong>the</strong> Andes orographic divide where <strong>the</strong> relationshipbetween precipitation and westerly w<strong>in</strong>d strength is positive(Fig. 1b), show a transition from a lake to a mire at that site(between 11.2 and 10.9 ka), <strong>in</strong> concert with an abrupt and susta<strong>in</strong>ed<strong>in</strong>crease <strong>in</strong> “hygrophyte” pollen taxa between 11 and 8 ka (Fig. 4b)that reflects <strong>the</strong> colonisation of a former lake by <strong>the</strong> pr<strong>in</strong>cipal mireform<strong>in</strong>g plant, Marsippospermum spp., and peatland dynamics<strong>the</strong>reafter (Fesq-Mart<strong>in</strong> et al., 2004), a process known as terrestrialisation,when lakes <strong>in</strong>fill under stable or dropp<strong>in</strong>g lake-levels(Futyma and Miller, 1986; Korhola, 1995; Brugam and McCanceJohnson, 1997; Weckstrom et al., 2010), imply<strong>in</strong>g a transition todrier climate between 11.2 and 10.9 ka. After w8.5 ka, an <strong>in</strong>crease<strong>in</strong> moisture is reflected by <strong>the</strong> establishment of Magellanic Moorlandtaxa (Astelia pumila and Donatia fascicularis) at <strong>the</strong> site (Fesq-Mart<strong>in</strong> et al., 2004).A partially published pollen record from Lago Tamar (52 54.21 0 S,73 48.07 0 W; Lamy et al., 2010), located west of <strong>the</strong> Andes, showsa marked and susta<strong>in</strong>ed <strong>in</strong>crease <strong>in</strong> Misodendron pollen, a hemiparasiteon Nothofagus trees, between 11.5 and 8.5 ka (Fig. 4c; Lamyet al., 2010). The Misodendron peak is temporally synchronous with<strong>the</strong> negative moisture anomalies identified at Lago Guanaco and LagoCondorito (Fig. 4e, f). Studies of modern pollen dispersal <strong>in</strong> Sou<strong>the</strong>rnPatagonia reveal greatest Misodendron pollen content at <strong>the</strong> relativelydry and open forest-steppe ecotone (Markgraf et al.,1981; Paezet al., 2001). Numerical analysis of modern pollen data from <strong>the</strong>region reveals that Misodendron pollen content is positively correlatedwith Nothofagus and o<strong>the</strong>r forest <strong>in</strong>dicators (Dale et al., 2010).Thus, while <strong>in</strong>terpretation of <strong>the</strong> Misodendron curve is complicatedby <strong>the</strong>se factors, it is likely to reflect a trend toward canopy openness,which is consistent with a trend toward drier conditions.4.2.3. Western Patagonia firePeak charcoal activity is evident between 12.5 and 9.5 ka <strong>in</strong> siteslocated between 40 and 55 S, where ra<strong>in</strong>fall is positively correlatedwith zonal w<strong>in</strong>d speed (Heusser, 2003; Haberle and Bennett, 2004;Huber et al., 2004; Moreno, 2004; Whitlock et al., 2007; Abarzúaand Moreno, 2008; Massaferro et al., 2009; Markgraf and Huber,2010). The regional charcoal curve for SSA (Power et al., 2008)captures this trend remarkably well (Fig. 4d) and displays a clearcorrelation to <strong>the</strong> palaeovegetation <strong>in</strong>dices from Lago Guanaco andLago Condorito (Fig. 4e, f). The charcoal data <strong>in</strong>dicate <strong>in</strong>creas<strong>in</strong>gcharcoal from 12.5 to 9.5 ka; decreas<strong>in</strong>g charcoal from 9.5 to 3.5 ka;<strong>in</strong>creas<strong>in</strong>g charcoal to 2.5 ka; and little change <strong>the</strong>reafter (Fig. 4b).4.2.4. SummaryHigh relative moisture <strong>in</strong> <strong>the</strong> northwest Patagonian palaeovegetation<strong>in</strong>dex and low sou<strong>the</strong>rn South American charcoalvalues between 14 and 12 ka imply strong westerly flow that isconsistent with dry conditions and low lake-levels east of <strong>the</strong>Andes Cordillera at Lago Cardiel. The available evidence <strong>in</strong>dicatesa multi-millennial decrease (<strong>in</strong>crease) <strong>in</strong> moisture (fire) <strong>in</strong> northwestPatagonia from w12.5 ka. In southwest Patagonia, <strong>in</strong>creasedmoisture <strong>in</strong> Lago Guanaco, cool-wet conditions <strong>in</strong> Gran Campo-2(Fesq-Mart<strong>in</strong> et al., 2004) and moist conditions at Isla de losEstados (54 50S; 64 40W; Ponce et al., 2011) <strong>in</strong>dicates an asymmetryof moisture changes north and south of <strong>the</strong> modern westerlycore that implies a southward shift of <strong>the</strong> SWW at this time. After11 ka, negative moisture trends at Lago Condorito and L. Guanaco,warmer and drier conditions at Isla de los Estados (Ponce et al.,2011) and terrestrialisation of <strong>the</strong> Gran Campo-2 site <strong>in</strong>dicatea decrease <strong>in</strong> SWW-derived moisture that persists until w8 ka,reveal<strong>in</strong>g negative moisture trends across <strong>the</strong> entire zone ofwesterly <strong>in</strong>fluence, coeval with peak fire activity <strong>in</strong> high-ra<strong>in</strong>fallregions throughout Patagonia. East of <strong>the</strong> Andes, above modernlake-level at Lago Cardiel between w11 and 8 ka (Fig. 4a) anda trend toward <strong>in</strong>creased relative moisture across sites <strong>in</strong> Argent<strong>in</strong>a(Manc<strong>in</strong>i et al., 2008) are consistent with decreased SWW <strong>in</strong>fluence,reveal<strong>in</strong>g synchronous and co-variable changes <strong>in</strong> westerlyw<strong>in</strong>d behaviour north and south of <strong>the</strong> core of <strong>the</strong> SWW (rang<strong>in</strong>gbetween 41 and 52 S) that led to decreased moisture west of <strong>the</strong>Andes and less <strong>in</strong>tense foehn w<strong>in</strong>ds and <strong>in</strong>creased <strong>in</strong>cursions ofAtlantic Ocean moisture to <strong>the</strong> east.Increas<strong>in</strong>g relative moisture <strong>in</strong> <strong>the</strong> west and a concomitantdecrease <strong>in</strong> sou<strong>the</strong>rn South American charcoal is evident afterw8 ka toward an apparent ‘moisture maximum’ <strong>in</strong> northwestPatagonia between 6 and 5 ka. This pattern is consistent with<strong>in</strong>creas<strong>in</strong>g moisture <strong>in</strong> southwest Patagonia (Fig. 4e, f) anddecreased lake-levels at Lago Cardiel (Fig. 4a). Moreover, <strong>the</strong>setrends are consistent with a stepwise <strong>in</strong>crease <strong>in</strong> effective precipitation<strong>in</strong> central Chile at 8.6 ka and 5.6 ka (Fig. 4f) and a trendtoward drier climate conditions <strong>in</strong> Argent<strong>in</strong>a, east of <strong>the</strong> AndesCordillera (Manc<strong>in</strong>i et al., 2008). These regionally consistent trendsimply that <strong>the</strong> westerlies <strong>in</strong>creased <strong>in</strong> strength across <strong>the</strong>ir entirezone of <strong>in</strong>fluence. Moisture <strong>in</strong>dices diverge after 5 ka, with a slightmulti-millennial decrease <strong>in</strong> relative moisture evident <strong>in</strong> northwestPatagonia (Fig. 4e), and multi-millennial moisture <strong>in</strong>creases <strong>in</strong>southwest Patagonia and central Chile (Fig. 4f, h). Interest<strong>in</strong>gly,a lake-level regression start<strong>in</strong>g >8.7 ka at Laguna Potrok Aike(Fig. 4g), east of <strong>the</strong> Andes <strong>in</strong> a zone of zero statistical correlationbetween westerly w<strong>in</strong>d speed and precipitation (Fig. 1b), followedby a transgression w7 ka occurs <strong>in</strong> concert with <strong>the</strong> Lago Guanacopalaeovegetation (moisture) <strong>in</strong>dex located at <strong>the</strong> same latitude(52 S), suggest<strong>in</strong>g that multi-millennial scale changes <strong>in</strong> moistureat Laguna Potrok Aike have varied with similar tim<strong>in</strong>g and directionas records from areas positively correlated with westerly w<strong>in</strong>dstrength dur<strong>in</strong>g <strong>the</strong> Holocene. Charcoal values <strong>in</strong>crease after 3.5 kaand rema<strong>in</strong> stable, as does <strong>the</strong> level of Lago Cardiel (Fig. 4a, d). Theeffective precipitation curve from Laguna Aculeo shows fur<strong>the</strong>r<strong>in</strong>creases at 3.2 ka and 2 ka, reflect<strong>in</strong>g an <strong>in</strong>crease <strong>in</strong> westerlyderived moisture <strong>in</strong> that region (Fig. 4h).5. Sou<strong>the</strong>rn Africa5.1. Present environmentThe climate of sou<strong>the</strong>rn Africa is dom<strong>in</strong>ated by <strong>the</strong> sub-tropicalhigh pressure system result<strong>in</strong>g from descend<strong>in</strong>g Hadley Cell airand <strong>the</strong> region is generally dry (Tyson and Preston-Whyte, 2000).Seasonal <strong>in</strong>trusions of monsoonal ra<strong>in</strong> are important dur<strong>in</strong>gsummer <strong>in</strong> <strong>the</strong> north of <strong>the</strong> region. The SWW exert a direct <strong>in</strong>fluenceon <strong>the</strong> climate of <strong>the</strong> southwest tip (<strong>the</strong> Western Cape), impart<strong>in</strong>ga w<strong>in</strong>ter wet Mediterranean climate <strong>in</strong> that region, while <strong>the</strong> EasternCape is reliant on easterly sourced precipitation delivered dur<strong>in</strong>g <strong>the</strong>summer months when westerly flow is weak and <strong>the</strong> trade w<strong>in</strong>dsare displaced southward (Tyson and Preston-Whyte, 2000). Thiseasterly precipitation source is bolstered by <strong>the</strong> warm southwardbound Agulhas Current that flows along <strong>the</strong> east African coast,result<strong>in</strong>g <strong>in</strong> frequent mists (Eeley et al., 1999). The dom<strong>in</strong>ance of <strong>the</strong>trade w<strong>in</strong>ds and <strong>the</strong> cold north-flow<strong>in</strong>g Benguela Current (driven <strong>in</strong>
40M.-S. Fletcher, P.I. Moreno / Quaternary International 253 (2012) 32e46part by <strong>the</strong> SWW) result <strong>in</strong> a hyper-arid desert on <strong>the</strong> western coastof sou<strong>the</strong>rn Africa (Namibia) that grades to <strong>the</strong> Mediterraneanclimate zone of <strong>the</strong> Western Cape fur<strong>the</strong>r south as <strong>the</strong> SWW becomeimportant (Tyson and Preston-Whyte, 2000).Sou<strong>the</strong>rn Africa hosts one of <strong>the</strong> most diverse vegetation typeson Earth, <strong>the</strong> Fynbos (Muc<strong>in</strong>a and Ru<strong>the</strong>rford, 2006). The Fynbos iscomprised of a startl<strong>in</strong>gly diverse array of shrubs that grow on <strong>the</strong>nutrient-poor geologies of <strong>the</strong> Western Cape Mediterraneanclimate zone (Muc<strong>in</strong>a and Ru<strong>the</strong>rford, 2006). To <strong>the</strong> north andcentre of sou<strong>the</strong>rn Africa, aridity prevails and desert-adapted florapredom<strong>in</strong>ate that grade <strong>in</strong> to savannah biomes as <strong>the</strong> monsoonal<strong>in</strong>fluence <strong>in</strong>creases. The higher relative moisture <strong>in</strong> <strong>the</strong> sou<strong>the</strong>astgives rise to a slightly more woody vegetation that transitions <strong>in</strong>to<strong>the</strong> grasslands and savannah of <strong>the</strong> nor<strong>the</strong>rn <strong>in</strong>terior and humidforest patches <strong>in</strong> relatively moist areas on <strong>the</strong> east coast (Eeleyet al., 1999; Muc<strong>in</strong>a and Ru<strong>the</strong>rford, 2006).5.2. Palaeoenvironmental records5.2.1. Western Cape organic sediment accumulationThe dry environment of sou<strong>the</strong>rn Africa is preclusive to organicsediment accumulation, a facet that would have been amplified(weakened) under past climate regimes <strong>in</strong> which drier (wetter)conditions prevailed. Preserved organic beds spann<strong>in</strong>g <strong>the</strong> latePleistocene to <strong>the</strong> present are rare <strong>in</strong> <strong>the</strong> Western Cape (32e34 S;Fig. 1a), a region wholly dependent on <strong>the</strong> SWW for precipitationthat displays a positive correlation between westerly w<strong>in</strong>d speedand precipitation <strong>in</strong> <strong>the</strong> modern climate (Fig. 1b). Fig. 5b showsa plot of <strong>the</strong> number published organic profiles from <strong>the</strong> WesternCape region at 1000-year time steps (based on <strong>the</strong> studies ofSchalke, 1973; Meadows, 1988; Meadows and Sugden, 1991; Street-Perrott and Perrott, 1993; Meadows et al., 1996; Meadows andBaxter, 2001). A very low number of profiles date to <strong>the</strong> periodbetween 14 and 9 ka. While this trend may reflect non-climaticfactors, such as a lack of discovery and/or publication of records,<strong>the</strong> long history of palaeoclimatic research <strong>in</strong> this region suggeststhat <strong>the</strong> lack of organic profiles between 14 and 9 ka may reflectaridity through all or <strong>the</strong> latter part of this time-span. After 9 ka, <strong>the</strong>number of published organic sequences <strong>in</strong>creases two-fold andrema<strong>in</strong>s high until <strong>the</strong> present, possibly reflect<strong>in</strong>g more humidconditions s<strong>in</strong>ce 9 ka.Fig. 5. Palaeoenvironmental data from sou<strong>the</strong>rn Africa plotted on a calendar age scale:(a) Matputaland pollen record Podocarpus values (Site 2, Fig. 1a; F<strong>in</strong>ch and Hill, 2008);(b) Western Cape organic accumulation (Site 1, Fig. 1a; Schalke, 1973; Meadows, 1988;Meadows and Sugden, 1991; Street-Perrott and Perrott, 1993; Meadows et al., 1996;Meadows and Baxter, 2001). Timescales were developed based on calendar yearswhenever <strong>the</strong> orig<strong>in</strong>al records were published <strong>in</strong> radiocarbon age scales. Radiocarbondates were calibrated us<strong>in</strong>g Calib 6.1 (Stuiver et al., 2010) and l<strong>in</strong>ear <strong>in</strong>terpolationsdeveloped between <strong>the</strong>se calibrated dates. See Fig. 1a for <strong>the</strong> location of sites. Greyshad<strong>in</strong>g <strong>in</strong>dicates <strong>the</strong> early Holocene (11e8 ka) period of weak westerly flow <strong>in</strong> <strong>the</strong>Sou<strong>the</strong>rn Hemisphere.5.2.2. Coastal sou<strong>the</strong>ast Africa e Matputaland vegetationA pollen record from Matputaland <strong>in</strong> coastal sou<strong>the</strong>ast Africa(28 S; Fig. 1a), with<strong>in</strong> <strong>the</strong> zone of negative correlation betweenwesterly w<strong>in</strong>d speed and precipitation (Fig. 1b) and a regiondependent on easterly <strong>in</strong>cursions of moisture from <strong>the</strong> IndianOcean, tracks moisture-driven vegetation changes through <strong>the</strong> lateQuaternary (F<strong>in</strong>ch and Hill, 2008). F<strong>in</strong>ch and Hill (2008) reconstructedrelative moisture changes at <strong>the</strong>ir Matputaland site basedpredom<strong>in</strong>antly on changes <strong>in</strong> Podocarpus pollen, a humid foresttree (Eeley et al., 1999) that is absent from <strong>the</strong> site today (F<strong>in</strong>ch andHill, 2008). Based on this premise <strong>the</strong>y <strong>in</strong>ferred relatively dryconditions and low Podocarpus pollen values (70%)between 11 and 9.2 ka, after which a steady decl<strong>in</strong>e <strong>in</strong> valuesbetween 9.2 and 5.4 ka <strong>in</strong>dicates a decl<strong>in</strong>e <strong>in</strong> available moisture anda reduction of this forest type (Fig. 5a). After 5.4 ka, decreas<strong>in</strong>g andvariable Podocarpus values possibly <strong>in</strong>dicate a variable climatetrend<strong>in</strong>g toward drier conditions (Fig. 5a).5.2.3. SummaryThe pattern of multi-millennial changes <strong>in</strong> moisture regimes <strong>in</strong>sou<strong>the</strong>rn South Africa is consistent with <strong>the</strong> modern relationshipbetween westerly flow and precipitation. Little <strong>in</strong>formation ispresent for conditions <strong>in</strong> <strong>the</strong> Western Cape region between 14 and11 ka, although a review of palaeoenvironmental data from thisregion depicts <strong>the</strong> LGM and late Pleistocene as cool and wet(Meadows and Baxter, 1999). Dry conditions (low Podocarpusvalues) on <strong>the</strong> sou<strong>the</strong>ast coast at this time are consistent with <strong>the</strong>modern relationship between zonal w<strong>in</strong>d strength and precipitationunder <strong>in</strong>creased westerly flow and <strong>the</strong> evidence suggestsa streng<strong>the</strong>n<strong>in</strong>g of westerly flow across sou<strong>the</strong>rn South Africabetween 14 and 11 ka. A sharp <strong>in</strong>crease <strong>in</strong> Podocarpus pollen and<strong>the</strong> establishment of humid forest on <strong>the</strong> sou<strong>the</strong>ast coast betweenw11 and 8 ka is consistent with decreased westerly flow anda concomitant <strong>in</strong>crease <strong>in</strong> <strong>in</strong>cursions of easterly moisture sourcesand coastal mists. The Western Cape is represented by very feworganic sequences between 11 and 9 ka, with <strong>the</strong> review ofMeadows and Baxter (1999) suggest<strong>in</strong>g dry conditions through thistime, and it is possible that weak SWW flow between 11 and 9 karesulted <strong>in</strong> enhanced aridity, desiccat<strong>in</strong>g any earlier organic deposition.From 9 ka onward, stronger westerly flow and moistconditions <strong>in</strong> <strong>the</strong> Western Cape region are suggested by an <strong>in</strong>crease<strong>in</strong> <strong>the</strong> number of published organic sequences and is consistentwith a review by Meadows and Baxter (1999), while a substantialdecl<strong>in</strong>e <strong>in</strong> Podocarpus pollen toward a low at 5.5 ka reflects dry<strong>in</strong>g<strong>in</strong> <strong>the</strong> east, a pattern consistent with <strong>the</strong> effects of stronger SWWon precipitation <strong>in</strong> <strong>the</strong> modern climate (Fig. 1b), although it must benoted that this latter region lies <strong>in</strong> a summer-ra<strong>in</strong>fall region,whereas westerly w<strong>in</strong>ds predom<strong>in</strong>antly <strong>in</strong>fluence w<strong>in</strong>ter wea<strong>the</strong>rpatterns <strong>in</strong> this region. The number of organic sequences changeslittle from 9 ka to <strong>the</strong> present <strong>in</strong> <strong>the</strong> Western Cape, while a markeddrop <strong>in</strong> Podocarpus at 3 ka followed by an <strong>in</strong>crease centred on 2 kareveals variability <strong>in</strong> moisture regime on <strong>the</strong> sou<strong>the</strong>ast coast thatmay be attributable to changes <strong>in</strong> westerly flow.6. Discussion6.1. The Sou<strong>the</strong>rn Westerly W<strong>in</strong>ds s<strong>in</strong>ce 14 kaThis syn<strong>the</strong>sis, analysis, and re<strong>in</strong>terpretation of selectedSou<strong>the</strong>rn Hemisphere palaeoenvironmental records represents <strong>the</strong>first attempt at <strong>in</strong>tegrat<strong>in</strong>g data from all Sou<strong>the</strong>rn Hemispherelandmasses relevant to westerly w<strong>in</strong>d changes <strong>in</strong> <strong>the</strong> post-LGMperiod. Despite <strong>the</strong> limited chronological control of many of <strong>the</strong>