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The Paleontological Society Special PublicationsA Publication of The Paleontological SocietySeries EditorRussell D. (“Tim”) WhiteDivision of Invertebrate PaleontologyPeabody Museum of Natural HistoryYale University170 Whitney Ave., P.O. Box 208118New Haven, CT 06520Copyright © 2002 by The Paleontological SocietyAll rights reserved. This entire publication may not be reproduced, stored in a retrievalsystem, or transmitted in any form or by any means, electronic, mechanical,photocopying, recording, or otherwise, without permission in written form from theSecretary of the Society. Diagrams, figures, tables, illustrations, and graphs may bereproduced by photocopying and distributing free of charge for educational purposes, ifproper credit is given to the author(s) and the Society.Printed by Yale University Reprographics & Imaging Services, New Haven, CT.