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PALEONTOLOGICAL SOCIETY PAPERS, V. 8, 2002WARÉN, A. 1981. Eulimid gastropods parasitic on echinoderms in the New Zealand region. New Zealand Journalof Zoology, 8:313–324.WARÉN, A., D. R. NORRIS, AND J. T. TEMPELADO. 1994. Description of four new eulimid gastropods parasitic onirregular sea urchins. Veliger, 37:141–154.WARÉN, A., AND M. R. CROSSLAND. 1991. Revision of Hypermastus Pilsbry, 1899 and Turveria Berry, 1956 (Gastropoda:Prosobranchia: Eulimidae), two genera parasitic on sand dollars. Records of Australian Museum, 43:85–112.WILTSE, W. I. 1980. Predation by juvenile Polinices duplicatus (Say) on Gemma gemma (Totton). Journal ofExperimental Marine Biology and Ecology, 42:187–199.WILSON, J. B. 1967. Palaeoecological studies on shell-beds and associated sediments in the Solway Firth. ScottishJournal of Geology, 3:329–371.WILSON, M. V. H. 1987. Predation as a source of fish fossils in Eocene lake sediments. Palaios, 2:497–504.WILSON, M. A., AND T. J. PALMER. 2001. Domiciles, not predatory borings: A simpler explanation of the holes inOrdovician shells analyzed by Kaplan and Baumiller, 2000. Palaios, 16:524–525.WOELKE, C. E. 1957. The flatworm Pseudostylochus ostreophagus Hyman, a predator of oysters. Proceedings ofthe National Shellfisheries Association, 47:62–67.WOOD, R. In press. Predation in ancient reef-builders. In P. H. Kelley, M. Kowalewski, and T. A. Hansen (eds.), Predator-Prey Interactions in the Fossil Record. Topics in Geobiology Series, Plenum Press/Kluwer, New York.WODINSKY, J. 1969. Penetration of the shell and feeding on gastropods by Octopus. American Zoologist, 9:997–1010.YOCHELSON, E. L., D. DOCKERY, AND H. WOLF. 1983. Predation on sub-Holocene scaphopod mollusks from southernLouisiana. United States Geological Survey Professional Paper, 1282:1–13.YONGE, C. M. 1964. Rock borers. Sea Frontiers, 10:106–116.YOUNG, D. K. 1969. Okadaia elegans, a tube-boring nudibranch mollusc from the central and west Pacific.American Zoologist, 9:903–907.ZAR, J. H. 1999. Biostatistical analysis, 4 th ed. Prentice Hall, Upper Saddle River, 663 p.ZIEGELMEIER, E. 1954. Beobachtungen über den Nahrungserwerb bei der Naticide Lunatia nitida Donovan(Gastropoda Prosobranchia). Helgoländer wissentschaftliche Meeresuntersuchungen, 5:1–33.ZILCH, A. 1959. Euthyneura, p. 1–701. In O. H. Schindewolf (ed.), Handbuch der Paläozoologie, 6. Berlin.ZOTNIK, M. 2001. Size-related changes in predatory behaviour of naticid gastropods from the middle MioceneKorytnica Clays, Poland. Acta Palaeontologica Polonica, 46:87–97.ZUSCHIN, M., AND R. J. STANTON, JR. 2001. Experimental measurement of shell strength and its taphonomicinterpretation. Palaios, 16:161–170.40
KOWALEWSKI—ANALYTICAL METHODSAPPENDIX 1—A SAS/IML and SAS/STAT programto test for non-random distribution of traces acrosstaxa. The program generates four outputs: (1)frequency of traces for each taxon and eachiteration; (2) Null proportion of taxa expected tocontain traces sorted by iteration; (3) Null highestper-taxon frequency of traces sorted by iteration;and (4) Null highest per-taxon frequency of tracesby iteration and taxon. Written by M. Kowalewski.________________________________________________%let prob=0.011; * - assemblage-level drilling frequency (AF);%let times=999; * - number of iterations;PROC IML;X={292,280,280,280,220,200,200,200,140,140,140,120,120,100,100,100,100,100,100,100,80,80,80,80,80,80,60,60,60,60,60,60,60,60,60,60,60};* - enter the specimen numbers for all taxa here;START drill(X,fin);Z=X; a=1-&prob; b=&prob; k=nrow(Z);DO i=1 to k;new=shape(i,Z[i],1);size=shape(Z[i],Z[i],1);newsize=new||size;new2=new2//newsize;END;out1=new2; k2=nrow(out1);DO j=1 to k2;c=rantbl(0,a,b); c1=c-1;c2=c2//c1;END;out2=c2; out3=out1||out2;fin=out3;FINISH drill;START simul(X,out);do i=1 to ×run drill(X,fin);fin2=fin||shape(i,nrow(fin),1);fin3=fin3//fin2;end;out=fin3;FINISH simul;RUN simul(X,out);CREATE new from out;APPEND from out;CLOSE new;RUN;DATA output;set new; taxon=col1; nspec=col2; drill=col3; iter=col4;PROC sort; by iter taxon nspec;PROC univariate noprint;by iter taxon nspec; var drill;output out=final n=n mean=mean sum=sum;title1*Simulation Output for &times iterat & drilling prob &prob*;title2’Total output’;PROC print;var iter taxon nspec mean sum;DATA new2;set final;if sum>0 then sum=1;PROC univariate noprint;by iter; var sum;output out=final2 mean=mean;title2’Proportion of taxa drilled by iteration’;PROC print;PROC univariate data=final noprint;by iter; var mean;output out=final3 max=max;title2’The highest per-taxon drilling frequency by iteration’;PROC print;PROC sort data=final; by iter nspec;PROC univariate noprint;by iter nspec; var mean;output out=final4 max=max;DATA new3;set final4; keep iter nspec max;title2’The highest drilling frequency by taxon sample size byiteration’;PROC print;QUIT;_______________________________________________________APPENDIX 2—A SAS/IML and SAS/STAT programfor a two-sample bootstrap test for difference inmeans for circular (azimuth) data. The algorithmfor the Watson-Williams two-sample test is basedon equations from Zar (1999, p. 626, Example27.8). F statistic is not corrected (i.e., K=1; seeZar, 1999, Equation 27.11). The correction is notneeded here because the probability densityfunction for F is established empirically throughbootstrap simulation. Bootstrap probability [p] iscalculated as follows: p=s+1/i+1, where s is thenumber of bootstrap values larger than or equal toactual F and i is the number of iterations (thisequation includes the actual samples in computingp; see Manly, 1995). Written by M. Kowalewski.Note: This code was used recently by Dietl andAlexander (2000) to analyze changes in siteselectivity of drilling on Cenozoic gastropods._______________________________________%let TIMES=999; *—number of times to randomize;%let DATA1=’ang1.dat’; *—file containing first variable;%let DATA2=’ang2.dat’; *—file containing second variable;Title1’2-sample bootstrap for circular data (Watson-Williamstest)’;Title2’written by Michal Kowalewski, October 11, 1999';DATA data1; infile &DATA1; input var1;DATA data2; infile &DATA2; input var2; RUN;PROC IML; %let pi=3.1415926535;USE data1; READ all var{var1} into X1;USE data2; READ all var{var2} into X2;START RANVEC(in,v_out); k=nrow(in); v_index=in;DO i=1 to k; rand=floor((k-i+1)*ranuni(0) + 1);v_ran=v_ran||v_index[rand]; v_index=remove(v_index,rand); END;v_out=v_ran; FINISH RANVEC;START MIXUP(X,times,template);n=nrow(X); template=t(1:n)*j(1,times,1);DO i=1 to times; run ranvec(template[,i],out);template[,i]=t(out); END; DO i=1 to n; runranvec(t(template[i,]),out); template[i,]=out; END; FINISH MIXUP;START WATSON(D1,D2,F);Y1=D1/(180/&pi); Y2=D2/(180/&pi); Y=Y1//Y2;C1=sum(COS(Y1))/nrow(Y1); S1=sum(SIN(Y1))/nrow(Y1);R1=sqrt(C1**2+S1**2)*nrow(Y1); a1=arcos(C1/(R1/nrow(Y1)))*(180/&pi);C2=sum(COS(Y2))/nrow(Y2); S2=sum(SIN(Y2))/nrow(Y2);R2=sqrt(C2**2+S2**2)*nrow(Y2); a2=arcos(C2/(R2/41
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Page 4 and 5: The Paleontological Society Special
Page 6 and 7: AuthorsRICHARD K. BAMBACHBotanical
Page 9 and 10: THE FOSSIL RECORD OF PREDATIONMicha
Page 13 and 14: INTRODUCTIONTHE FOSSIL RECORD OF PR
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