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The State of Circumpolar Walrus Populations

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from Clavering Island to the northern border <strong>of</strong> the Northeast Water yielded a fully corrected<br />

summer estimate <strong>of</strong> 1,430 (CV = 0.45) walruses (Born et al. 2009). Modelling suggests this<br />

population has recovered from its historical depletion but the trajectory <strong>of</strong> this recovery is<br />

uncertain due to lack <strong>of</strong> a population specific growth rate estimate (Witting and Born 2014).<br />

Svalbard – Franz Josef Land Population (S-FJL)<br />

<strong>The</strong> Norwegian archipelago <strong>of</strong> Svalbard and Russian archipelago <strong>of</strong> Franz Josef Land share a<br />

walrus population (Wiig et al. 2014; Figure 4). <strong>The</strong>se animals are genetically similar (Andersen et<br />

al. 1998; Born et al. 2001), move between the archipelagos (Wiig et al. 1996; Freitas et al. 2009;<br />

Hamilton et al. 2015), and are separated from other walruses by wide distributional gaps (Born<br />

1984; Born and Gjertz 1993; Gjertz and Wiig 1994). In 2010 an adult male that was genetically<br />

similar to walruses in this population travelled from the Faroe Islands to Svalbard, a distance <strong>of</strong><br />

2216 km between 29 March and 25 April (Born et al. 2014). <strong>The</strong> relationship <strong>of</strong> Atlantic walruses<br />

in this population to those in the Kara Sea - southern Barents Sea –Novaya Zemlya population is<br />

uncertain (Born et al. 1995; NAMMCO 2006; Boltunov et al. 2010; Shitova et al. 2014b).<br />

<strong>Walrus</strong>es in Svalbard follow the same seasonal migration pattern regardless <strong>of</strong> annual variations<br />

in ice and temperature regimes (Freitas et al. 2009). Summer habitat use for both sexes appears<br />

to be driven by feeding requirements and the availability <strong>of</strong> haulouts or sea ice. Most <strong>of</strong> the males<br />

summer in Svalbard and most <strong>of</strong> the females and calves remain in northeastern parts <strong>of</strong> Svalbard,<br />

in the Franz Josef Land archipelago, or between them on Victoria Island (Lydersen et al. 2008;<br />

Gavrilo 2010; Kovacs et al. 2014; Wiig et al. 2014). To reach breeding areas the males actively<br />

travel through areas <strong>of</strong> dense ice cover towards Franz Josef Land in winter, regardless <strong>of</strong> sea ice<br />

advances and retreats (Freitas et al. 2009).<br />

<strong>Walrus</strong>es in Svalbard were protected from harvesting in 1952 after having been brought to the<br />

brink <strong>of</strong> extinction by 350 years <strong>of</strong> unregulated removals (Anon. 1952; Kovacs et al. 2014). Prior<br />

to commercial hunting the population must have been very large (Reeves 1978; Gjertz et al. 1998;<br />

Weslawski et al. 2000). During the first 30 years <strong>of</strong> protection, about 100 animals became<br />

established within the archipelago. <strong>The</strong>y are presumed to have come from Franz Josef Land to the<br />

east. Since then a marked recovery has occurred in the abundance <strong>of</strong> walruses in Svalbard.<br />

Systematic surveys that covered all current and historical haulout sites were flown in August 2006<br />

(Lydersen et al. 2008) and late July to mid-August 2012 (Kovacs et al. 2014). <strong>The</strong> surveys,<br />

adjusted to account for animals that were in the water, estimated there were 2,629 (95% CI =<br />

2,318 – 2,998) walruses in the Svalbard area in 2006 and 3,886 (95% CI = 3,553-4,262) in 2012.<br />

This represents an average annual increase <strong>of</strong> nearly 8% (Kovacs et al. 2014), which matches the<br />

theoretical maximum rate <strong>of</strong> growth that has been calculated for recovering walrus populations

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