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A Guide for Terrestrial Gastropod Identification - University of ...

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Helicodiscus nummus (Vanatta, 1899) G1G2 AR, KY, OK, TX<br />

AL, AR, CT, DE, FL, GA, IA, IL, IN, KS, KY, LA, MA,<br />

MD, ME, MI, MN, MO, MS, NC, NE, NH, NJ, NY, OH,<br />

OK, PA, RI, SC, TN, TX, VA, VT, WI, WV; Canada: NB,<br />

Helicodiscus parallelus (Say, 1817) G5<br />

NS, ON, QC, NF<br />

Helicodiscus punctatellus Morrison, 1942 G1 KY, TN<br />

Helicodiscus salmonaceus Hemphill, 1890 G2 ID, WA<br />

Helicodiscus saludensis (Morrison, 1937) G1 NC, SC<br />

CT, IA, IL, IN, KY, MA, MD, ME, MI, MN, NH, NY, PA,<br />

Helicodiscus shimeki Hubricht, 1962 G4G5<br />

TX, VA, VT, WI, WV; Canada: ON<br />

AL, AR, CA, DE, IL, IN, KS, KY, LA, MD, MI, MO, NE,<br />

NJ, NM, NY, OH, OK, PA, TN, TX, VA, WI, WY; Canada:<br />

Helicodiscus singleyanus (Pilsbry, 1889) G5<br />

ON<br />

Helicodiscus tridens (Morrison, 1935) G2 OK, TX<br />

Helicodiscus triodus Hubricht, 1958 G2 NC, VA, WV<br />

Polygyriscus virginianus (P.R. Burch, 1947) G1 LE VA<br />

Helminthoglyptidae<br />

Mark A. Ports, Great Basin College<br />

The family Helminthoglyptidae is a diverse and widespread group <strong>of</strong> land snails from<br />

Cali<strong>for</strong>nia, Oregon, Arizona, New Mexico, and west Texas. This family is composed <strong>of</strong> eleven<br />

genera with approximately 83 species <strong>of</strong> Helminthoglypta, 13 species <strong>of</strong> Sonorelix, 9 species <strong>of</strong><br />

Micrarionta, 1 species <strong>of</strong> Chamaeariontales, 1 species <strong>of</strong> Herpeterous, 8 species <strong>of</strong> Xerarionta,<br />

2 species <strong>of</strong> Cahuillus, 7 species <strong>of</strong> Eremarionta, 1 species <strong>of</strong> Noyo, 4 species <strong>of</strong> Rothelix, and<br />

72 species <strong>of</strong> Sonorella.<br />

The Helminthoglyptidae vary in diameter from 5 mm to 4 cm, and range from a dark<br />

brown color to a pearly white color. The shells are generally depressed-helicoid in shape, thin in<br />

texture, with a thin, reflexed peristome, normally without aperatural teeth. Most have a single<br />

dark supra-peripheral band while some <strong>of</strong> the Mohave Desert species may not have any band. In<br />

all <strong>of</strong> these genera (especially the Sonorella) the best way to identify them is the dissection <strong>of</strong><br />

their genitalia or the study <strong>of</strong> their phylogeny.<br />

This family occupies habitat types that include wet temperate <strong>for</strong>est in Oregon and<br />

northern Cali<strong>for</strong>nia, dry coniferous and deciduous <strong>for</strong>ests <strong>of</strong> the west side <strong>of</strong> the Sierra Nevada,<br />

coastal mountains and canyons <strong>of</strong> conifers and oak, chaparral brush <strong>of</strong> coastal Cali<strong>for</strong>nia, desert<br />

valleys and mountains <strong>of</strong> Cali<strong>for</strong>nia, Arizona, Texas and New Mexico. Most species <strong>of</strong> this<br />

family are associated with rocky or talus slopes and ledges <strong>of</strong> dolomite, granite, or shale.<br />

Existing in large to small colonies most species are active in the morning and evening or after<br />

31

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