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A Guide for Terrestrial Gastropod Identification - University of ...

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Holospira danielsi Pilsbry and Ferriss, 1915 G3G4 AZ, TX<br />

Holospira ferrissi Pilsbry, 1905 G2 AZ<br />

Holospira goldfussi (Menke, 1847) G2G3 TX<br />

Holospira hamiltoni Dall, 1897 G1 TX<br />

Holospira mesolia Pilsbry, 1912 G1 TX<br />

Holospira metcalfi F.G. Thompson, 1974 G1 NM<br />

Holospira millestriata Pilsbry and Ferriss, 1915 G1G2Q AZ<br />

Holospira montivaga Pilsbry, 1946 G2 AZ, NM, TX<br />

Holospira oritis Pilsbry and Cheatum, 1951 G1 TX<br />

Holospira pasonis Dall, 1895 G1 TX<br />

Holospira pityis Pilsbry and Cheatum, 1951 G1 TX<br />

Holospira riograndensis Pilsbry, 1946 G1 TX<br />

Holospira sherbrookei Gilbertson, 1989 G1 AZ<br />

Holospira tantalus Bartsch, 1906 G1G2 AZ<br />

Holospira whetstonensis Pilsbry and Ferriss, 1923 G1G2 AZ, NM<br />

Holospira yucatanensis Bartsch, 1906 G1 TX<br />

Metastoma roemeri (Pfeiffer, 1848) G4 NM, TX<br />

Microceramus pontificus (Gould, 1848) G2G3 FL<br />

Microceramus texanus (Pilsbry, 1898) G2 TX<br />

Valloniidae<br />

Jochen Gerber, Field Museum <strong>of</strong> Natural History<br />

The valloniids are minute snails (ca. 1.5-4 mm in diameter) <strong>of</strong> uni<strong>for</strong>mly white, greyish<br />

or brown shell color. Their shells are mostly depressed-helicoid, but some species are more<br />

globular. Many species have an ornamentation <strong>of</strong> transverse ribs and sometimes spiral lines.<br />

The aperture is either simple (Planogyra, Zoogenetes) or expanded (Vallonia). In some Vallonia<br />

species there is a threshold-like callus within the aperture, however, all valloniids lack apertural<br />

teeth or lamellae. The umbilicus is wide to very wide, with the exception <strong>of</strong> the tall-helicoid<br />

Zoogenetes which has a merely per<strong>for</strong>ate shell.<br />

The distribution <strong>of</strong> the family is holarctic. In North America they have a mainly northern<br />

distribution, with some Vallonia species extending further southward in the Western mountain<br />

ranges and into Northern Mexico. Some Vallonia species (V. costata, V. excentrica, V. pulchella)<br />

have been accidentally introduced in many areas outside their natural range.<br />

Vallonia species live usually in open habitats, such as meadows, lawns, and on exposed<br />

rock outcrops, occasionally extending into more open <strong>for</strong>est vegetation, but they avoid dense<br />

woods. Moisture requirements differ by species. The two Planogyra species live in leaf litter in<br />

moist <strong>for</strong>ests and swampy areas. Zoogenetes is a snail <strong>of</strong> boreal <strong>for</strong>ests <strong>of</strong> NE North America<br />

and <strong>of</strong> high-mountain <strong>for</strong>est habitats in the Rockies.<br />

59

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