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A Guide for Terrestrial Gastropod Identification - University of ...

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Urocoptidae<br />

Kathryn E. Perez, Duke <strong>University</strong><br />

Urocoptidae is a large family related to the Bulimulidae and Achatinidae. The shells <strong>of</strong> snails in<br />

this family tend to be elongate with a tapering spire and an expanded apertural lip. The two<br />

genera (and species) found in Florida are similarly elongate, but Cochlodinella poeyana is<br />

smaller with wider whorls and typically has a decollate spire in contrast with Microceramus<br />

pontificus which is somewhat larger with a wider shell. The shape <strong>of</strong> the aperture is also very<br />

different.<br />

The majority <strong>of</strong> U.S. Urocoptids are in the genus Holospira. These are snails <strong>of</strong> the<br />

limestone terrain <strong>of</strong> the arid southwestern U.S. Holospira and related genera found in Mexico are<br />

very species-rich. Important characteristics <strong>for</strong> identification <strong>of</strong> Holospira species are the length<br />

and width <strong>of</strong> the shell, the number <strong>of</strong> whorls, the smoothness or ribbing <strong>of</strong> the shell, and the<br />

shape (angulate, ovate, or oblong) and folding <strong>of</strong> the aperture. For final identification the shell<br />

must be opened, usually by cutting or grinding away the penultimate whorl to look at the central<br />

axis <strong>of</strong> the shell. Lamellae are associated with the central axis <strong>of</strong> the shell, especially in the final<br />

whorl. These lamellae are variously developed and shaped in different species.<br />

Species <strong>of</strong> Holospira tend to live in isolated large colonies and be very restricted in<br />

geographic location, <strong>for</strong> example, found only in single mountain ranges. One exception to this is<br />

Metastoma roemeri which is found throughout central Texas to the Rio Grande Valley <strong>of</strong> New<br />

Mexico. It is most abundant around New Braunfels, Texas. Other Holospira species are found<br />

hanging attached to or under limestone ledges, a few species in West Texas are associated with<br />

Selaginella (Resurrection plant). They can also be found along canyon walls, under stones, dead<br />

Yucca stems, dead stems <strong>of</strong> Sotol and in limestone talus (Metcalf & Smartt, 1997). Specific<br />

localities and habitat discussions <strong>of</strong> each New Mexico Holospira species can be found in Land<br />

Snails <strong>of</strong> New Mexico (Metcalf & Smartt, 1997).<br />

TAXON AUTHOR<br />

G-<br />

RANK DISTRIBUTION<br />

Cochlodinella poeyana (d'Orbigny, 1841) G1G2 FL<br />

Coelostemma pyrgonasta F. G. Thompson, 1989<br />

Gilbertson and Worthington,<br />

G1 NM<br />

Holospira animasensis<br />

2003 G1G2 NM<br />

Holospira arizonensis Stearns, 1890 G2 AZ<br />

Holospira campestris Pilsbry and Ferriss, 1915 G3Q AZ<br />

Holospira chiricahuana Pilsbry, 1905 G2G3 AZ<br />

Holospira cionella Pilsbry, 1905 G3Q AZ<br />

Holospira cockerelli Dall, 1897 G1 NM<br />

Holospira crossei Dall, 1895 G2 NM<br />

58

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