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Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...

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E. Lukševičs, I. Zupiņš. Sedimentology, fa<strong>un</strong>a, <strong>and</strong> taphonomy of Pav āri<br />

103<br />

Naum., Leiotriletes immanus Isch., Dictyotriletes sp. have been identified by M.<br />

Stanichnikova from clayey interbeds (Savvaitova 1977). V. Talimaa has reported remains<br />

of following vertebrate taxa from the Nīgr<strong>and</strong>e Member in the territory of Lithuania:<br />

Chelyophorus sp., P<strong>and</strong>erichthys sp., Rhizodontidae gen. indet., Osteolepis? sp.,<br />

Onychodus sp., Dipterus? sp., Palaeoniscidae gen. indet. in the lower part of the member;<br />

Bothriolepis sp., Acanthodei gen. indet., Sarcopterygii gen. indet., P<strong>and</strong>erichthys sp.,<br />

Holoptychius sp. in the upper part (Žeiba <strong>and</strong> Savvaitova 1981). In this case the<br />

identification of some taxa also causes doubt, as identifiable remains of the genus<br />

P<strong>and</strong>erichthys so far have been recorded only from the Givetian – Frasnian interval in<br />

the deposits of the Gauja, Lode, Amata, <strong>and</strong> Dubniki Formations, while the genus<br />

Osteolepis is known mainly from the Middle Devonian.<br />

The eroded surface of the clay <strong>and</strong> clayey-silty rocks of the Nīgr<strong>and</strong>e Member is<br />

overlaid by the yellowish grey fine-grained s<strong>and</strong>stone <strong>and</strong> s<strong>and</strong> of the Pavāri Member.<br />

In most of the distribution area s<strong>and</strong> <strong>and</strong> s<strong>and</strong>stones consist of quartz with admixture of<br />

feldspars, <strong>and</strong> in some places contain great amo<strong>un</strong>t of mica. In some levels s<strong>and</strong>stone is<br />

well cemented by dolomite. The content of fine-grained s<strong>and</strong> increases in the sections<br />

towards the northern part of distribution area (boreholes 27, 1-Priekule, 15-Sab-<br />

Nākotne); towards the north-western margin the amo<strong>un</strong>t of the medium sized grains<br />

also increases. The facies of the upper part of the Pavāri Member is more variable,<br />

being composed of thin beds of clay, clayey siltstone, dolomite marl <strong>and</strong> feldsparquartzitic<br />

micaceous s<strong>and</strong>. The total thickness of the section is 16-21 meters. S<strong>and</strong> <strong>and</strong><br />

s<strong>and</strong>stones of the Pavāri Member contain a rich assemblage of fishes <strong>and</strong> tetrapods, the<br />

full content of which is described further below.<br />

S<strong>and</strong>stones of the upper Varkaļi Member overlie the eroded surface of the Pavāri<br />

Member. The lower part of the Varkaļi Member is composed of light greenish-grey<br />

very fine <strong>and</strong> fine-grained feldspar-quartz s<strong>and</strong>s <strong>and</strong> weakly cemented s<strong>and</strong>stones,<br />

frequently containing large amo<strong>un</strong>ts of mica. Parallel cross-bedding is characteristic<br />

for the s<strong>and</strong>stones. In some beds, smooth horizontal bedding is accentuated by the<br />

distribution of muscovite scales, fish bones <strong>and</strong> their fragments, <strong>and</strong> clay pebbles. The<br />

lower part of the Varkaļi Member crops out on the right bank of the Venta River near<br />

the former Ketleri hamlet. The conglomerate in the fine grained s<strong>and</strong>stones of the lower<br />

part of this member is formed of well ro<strong>un</strong>ded blue-grey pebbles of microcrystalline<br />

dolomite marl of usually small, but sometimes quite large (up to 5 – 7 cm) size, dolomite<br />

marl interlayers <strong>and</strong> vertebrate remains. Mainly small remains of fishes <strong>and</strong> tetrapods<br />

as well as ro<strong>un</strong>ded <strong>and</strong> abraded fragments of larger bones are to be fo<strong>un</strong>d in s<strong>and</strong>y<br />

rocks. During several excavations organized by D. Obruchev (1957), L. Lyarskaya<br />

(1970, 1973) <strong>and</strong> E. Lukševičs (1984, 1988) rather rich material of fossil vertebrates<br />

has been collected at the locality: fragments of the tr<strong>un</strong>k armour <strong>and</strong> disarticulated<br />

bones of the head shield of placoderm Bothriolepis ciecere, acanthodian spines, scales<br />

of “Devononchus” ketleriensis Gross <strong>un</strong> “D.“ tenuispinus (Gross), as well as scales of<br />

at least yet another acanthodian, scales <strong>and</strong> rare disarticulated bones of the head of the<br />

sarcopterygians Holoptychius cf. nobilissimus, Ventalepis ketleriensis Schultze,<br />

Glyptopomus? bystrowi (Gross), Cryptolepis grossi Vorobyeva, Platycephalichthys sp.,<br />

parasphenoideum of a l<strong>un</strong>gfish Orlovichthys cf. limnatis Krupina, remains of tetrapods<br />

Ventastega curonica Ahlberg, Lukševičs et Lebedev <strong>and</strong> Tetrapoda gen. indet. (Ahlberg<br />

et al. 1994). Comparison of the vertebrate assemblages from Ketleri <strong>and</strong> Pavāri sites

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