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Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...

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E. Lukševičs, I. Zupiņš. Sedimentology, fa<strong>un</strong>a, <strong>and</strong> taphonomy of Pav āri<br />

107<br />

Fig. 5. Bipolar orientation of the cross-bedding of the<br />

infilling s<strong>and</strong>stone (layer No. 6). N = 30.<br />

of skulls have also been fo<strong>un</strong>d, particularly in slump depressions or close to slumping<br />

structures. Fossils are rarely fo<strong>un</strong>d close to the margins of the supposed erosional canal;<br />

mainly scales of sarcopterygians have been collected there. Fish <strong>and</strong> tetrapod remains<br />

are usually brown or dark brown, rather soft <strong>and</strong> fragile, therefore many specimens<br />

kept in the museum collection bear cracks, as the results of breakage during transportation<br />

in the plaster jackets or during preparation.<br />

Fossil assemblage. Of the more than 1000 specimens collected from the Pavāri site,<br />

909 have been identified to the generic/species level. About 800 specimens are housed<br />

at the LDM. Identified taxa from the Pavāri site are listed in the Table 1, <strong>and</strong> persentages<br />

of each taxon are shown in the Table 2 <strong>and</strong> Figure 6.<br />

Taxonomic representation. The fossils assemblage from the Pavāri site seems to be<br />

taxonomically incomplete, lacking the remains of arthrodires, ptyctodonts <strong>and</strong><br />

chondrichthyans. Actinopterygian remains are extremely rare, including only one small<br />

incomplete scale ever fo<strong>un</strong>d. Arthrodires are known from the <strong>un</strong>derlying beds of the<br />

Žagare Formation, while ptyctodonts <strong>and</strong> actinopterygians have been fo<strong>un</strong>d in the<br />

Nīgr<strong>and</strong>e Member of the Ketleri Formation (Gross 1933, Lyarskaya <strong>and</strong> Savvaitova<br />

1974). Chondrichthyan remains are in general rare in Baltic Devonian deposits (Ivanov<br />

<strong>and</strong> Lukševičs 1994; Mark-Kurik <strong>and</strong> Karatajūte-Talimaa 2004). From the Famennian<br />

deposits they have so far been fo<strong>un</strong>d only in dolostones of the Kursa Formation <strong>and</strong><br />

s<strong>and</strong>stones of the Mūri <strong>and</strong> Tērvete formation (both are older than the Ketleri Formation).<br />

Underrepresentation of small fish or small skeletal parts of fish bodies are common<br />

features of fossil assemblages <strong>and</strong> is usually interpreted as a combination of biotic <strong>and</strong><br />

abiotic factors that destroy or remove small, delicate bones (Behrensmeyer 1991).<br />

Possibly the absence of ptyctodonts, chondrichthyans <strong>and</strong> actinopterygians in the material<br />

from Pavāri can be explained by the small size of the armour plates of ptyctodonts,<br />

scales <strong>and</strong> teeth of chondrichthyans <strong>and</strong> actinopterygian scales, although the possibility<br />

that this part of the Baltic sedimentary basin was not suitable for the existence of these<br />

fishes during the Ketleri time can not be excluded. Considering the presence of<br />

acanthodian scales one should note their much smaller size comparable with that of the<br />

s<strong>and</strong> grains.<br />

Material from the Natural History Museum of Latvia (collection No LDM G 81),<br />

collected in 1970, 1973 <strong>and</strong> 1995, has been used for comparison of the percentage of

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