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Zemes un vides zinātnes Earth and Environment Sciences - Latvijas ...

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C.G.Miller, T. Marss, H. Blom. New anaspid material from Britain <strong>and</strong> Estonia<br />

53<br />

(Fig. 1F) to more elongate in the holotype (Fig. 1C) <strong>and</strong> the Estonian material certainly<br />

falls within these variations. However, we prefer to classify the Estonian material <strong>un</strong>der<br />

T. concatenata gen. et sp. nov. which has only one row of tubercles per scale, <strong>and</strong> three<br />

of the six scales are saddle shaped in cross section (Fig. 2E), whereas T. j<strong>un</strong>cta gen. et<br />

sp. nov. scales show two to four rows of tubercles joined by more prominent ridges<br />

(Figs 1, 2C). The Ludlow age of T. concatenata gen. et sp. nov. suggests that the range<br />

of the genus Trimpleylepis should be Ludlow-Prídolí.<br />

Family RHYNCHOLEPIDIDAE Kiær, 1924<br />

Silmalepis cf. erinacea Blom, Märss <strong>and</strong> Miller, 2003<br />

Fig. 3G-H<br />

Material. One broken scale from Man Brook from uppermost Prídolí, Upper Silurian,<br />

Man Brook, near Trimpley, Shropshire, U.K.<br />

Description. Scale with wide smooth overlapped area; groove extends entire length of<br />

scale close <strong>and</strong> parallel to the sculpture margin on the main scale area. External sculpture<br />

of closely spaced fine tubercles of variable diameter followed by relatively closely<br />

spaced ridges that point into the gap between the following row. The ends of ridges are<br />

broken in this specimen but three distinct rows of ridges can be seen across the scale.<br />

Visceral side with single rib <strong>and</strong> no evidence of vascular canal openings. No histological<br />

sections have been made due to lack of material.<br />

Remarks. Precise identification of this scale is not possible due to lack of material <strong>and</strong> the<br />

preservation of the scale, which has broken spine-like ridges. Septentrionia mucronata<br />

Blom, Märss <strong>and</strong> Miller, 2002 has spine-like ridges that point between the next set of<br />

ridges, as the spines do in the material figured here. S. mucronata is also similar as it has<br />

a wide overlapped area, but it differs in having a narrow well developed ridge with granular<br />

sculpture (Blom et al. 2003, fig. 45g-h) in the position where there is a groove developed<br />

in the scale illustrated here. In our opinion the scale figured here (Fig. 3G, H) is much closer<br />

to Silmalepis erinacea Blom, Märss <strong>and</strong> Miller, 2002 that also has a coarse tuberculated<br />

to granular sculpture to the anterior of the main area of the scale (Blom et al. 2002, fig. 34a).<br />

The long spines inclined at a low angle to the main scale area of Silmalepis (Blom et al.<br />

2003, fig. 34b) look to have broken off in the material figured here. The broken spines also<br />

have a concave upper surface as they do in S. erinacea. As the spines are broken, <strong>and</strong><br />

because the scale figured here has a much broader overlapped area than typical S. erinacea,<br />

we consider the material certainly belongs within Silmalepis but identify S. erinacea with<br />

doubt. The material figured here extends the range of Silmalepis from mid Ludlow (Blom<br />

et al. 2003, fig. 10) to mid Ludlow-late Prídolí.<br />

RHYNCHOLEPIDIDAE gen. et sp. indet.<br />

Figs 2B, D; 3I-L<br />

Material. One intact scale, one small fragment, <strong>and</strong> two fragments that were used for<br />

thin sectioning, all from from Gardner’s Bank.<br />

Description. Scales with a narrow furrowed overlapped area that is incompletely<br />

preserved on most of specimens available. Low ridges on the main scale area are wellspaced<br />

with the ends nearest the anterior overlapped area slightly curved. Each ridge

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